Polystichum imbricans |
Polystichum kwakiutlii |
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imbricate sword-fern, narrow-leaf sword fern, rock sword-fern |
kwakiutl's hollyfern |
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Stems | ascending to erect. |
unknown. |
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Leaves | erect to arching back at tip, 2–8 dm; bulblets absent. |
(only distal portion known) with bulblets present. |
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Petiole | 1/4–1/3 length of leaf; scales abruptly diminishing in size distally and falling off early but retaining conspicuous tuft of brown scales at base. |
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Blade | linear-lanceolate to linear, 1-pinnate, base not or slightly narrowed. |
lanceolate, 2-pinnate, base probably narrowed. |
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Pinnae | oblong, slenderly lanceolate, or falcate, usually overlapping, in 1 plane or twisted out of plane of blade, 2–4 cm; base oblique, auricles well developed; margins serrulate-spiny with teeth ascending; apex cuspidate or apiculate with subapical teeth smaller than apical tooth; microscales lanceolate to linear with straight or sharply angular projections, sparse, on abaxial surface only. |
narrowly lanceolate, ca. 2–7 cm, base truncate to oblique, acroscopic proximal pinnule enlarged, apex obtuse. |
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Indusia | entire to sharply dentate. |
entire. |
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Spores | dark brown. |
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Pinnules | short-stalked, ovate-rhombic, acroscopic auricle ± well developed, margins finely spiny-dentate; apex acuminate; microscales filiform, lacking projections, dense abaxially, sparse adaxially. |
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Polystichum imbricans |
Polystichum kwakiutlii |
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Habitat | Habitat unknown. | |||||
Distribution |
CA; OR; WA; BC; only in the flora
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BC; Of conservation concern |
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Discussion | Polystichum imbricans is one of the postulated ancestors of two allopolyploids, P. californicum and P. scopulinum (D. H. Wagner 1979). Relationships to P. munitum are discussed under that species. Subspecies 2. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Polystichum kwakiutlii is known only from the type specimen, collected at Alice Arm, British Columbia (whether referring to inlet or town is unknown). This species is presumed to be one of the diploid progenitors of P. andersonii. It should be sought among the boreal 2-pinnate polystichums, from which it can be distinguished by the presence of bulblets. Polystichum kwakiutlii differs from P. andersonii in its completely divided pinnae and entire indusia. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 2. | FNA vol. 2. | ||||
Parent taxa | Dryopteridaceae > Polystichum | Dryopteridaceae > Polystichum | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Aspidium munitum var. imbricans, P. munitum subsp. imbricans, P. munitum var. imbricans | |||||
Name authority | (D. C. Eaton) D. H. Wagner: Pteridologia 1: 50. (1979) | D. H. Wagner: Amer. Fern J. 80: 50, fig. 1. (1990) | ||||
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