Polystichum imbricans |
Polystichum kruckebergii |
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imbricate sword-fern, narrow-leaf sword fern, rock sword-fern |
holly fern, Kruckeberg's holly fern, Kruckeberg's sword fern, Kruckeberg's sword fern fern |
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Stems | ascending to erect. |
ascending. |
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Leaves | erect to arching back at tip, 2–8 dm; bulblets absent. |
erect, 1–2.5 dm; bulblets absent. |
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Petiole | 1/4–1/3 length of leaf; scales abruptly diminishing in size distally and falling off early but retaining conspicuous tuft of brown scales at base. |
1/10–1/5 length of leaf, sparsely scaly; scales light brown, gradually diminishing in size distally. |
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Blade | linear-lanceolate to linear, 1-pinnate, base not or slightly narrowed. |
linear, 1-pinnate-pinnatifid, base narrowed. |
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Pinnae | oblong, slenderly lanceolate, or falcate, usually overlapping, in 1 plane or twisted out of plane of blade, 2–4 cm; base oblique, auricles well developed; margins serrulate-spiny with teeth ascending; apex cuspidate or apiculate with subapical teeth smaller than apical tooth; microscales lanceolate to linear with straight or sharply angular projections, sparse, on abaxial surface only. |
rhombic-ovate to short-falcate, proximal pinnae ± triangular; pinnae overlapping, twisted somewhat out of plane of blade, 0.5–1.5 cm; base oblique, acroscopic auricle well developed; margins shallowly incised to merely dentate or serrulate, teeth spreading and spiny at tip; apex acute with subapical and apical teeth same size; microscales lanceolate with few projections, confined to costa, on abaxial surface only. |
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Indusia | entire to sharply dentate. |
entire. |
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Spores | dark brown. |
dark brown. |
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2n | = 164. |
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Polystichum imbricans |
Polystichum kruckebergii |
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Habitat | Rocks and cliffs in subalpine to alpine habitats | |||||
Elevation | 1500–3200 m (4900–10500 ft) | |||||
Distribution |
CA; OR; WA; BC; only in the flora
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CA; ID; MT; NV; OR; UT; WA; BC
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Discussion | Polystichum imbricans is one of the postulated ancestors of two allopolyploids, P. californicum and P. scopulinum (D. H. Wagner 1979). Relationships to P. munitum are discussed under that species. Subspecies 2. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Polystichum kruckebergii is widely but sporadically distributed in small numbers in both the Sierra-Cascade and Rocky Mountain systems. Populations sometimes consist of only two or three dwarfed plants that are difficult to distinguish from P. scopulinum, with which they may occur. The spreading teeth of equal size at the pinna apex will usually distinguish this species. Polystichum kruckebergii is a tetraploid presumed to be of hybrid origin, with P. lonchitis and P. lemmonii as its diploid progenitors (W. H. Wagner Jr. 1973), although this hypothesis has not been confirmed. The hybrid with P. munitum has been found in Washington (P. S. Soltis et al. 1987) with both parents, and it is distinguished by intermediate morphology and abortive sporangia. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 2. | FNA vol. 2. | ||||
Parent taxa | Dryopteridaceae > Polystichum | Dryopteridaceae > Polystichum | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Aspidium munitum var. imbricans, P. munitum subsp. imbricans, P. munitum var. imbricans | |||||
Name authority | (D. C. Eaton) D. H. Wagner: Pteridologia 1: 50. (1979) | W. H. Wagner: Amer. Fern J. 56: 4. (1966) | ||||
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