Polystichum imbricans |
Polystichum dudleyi |
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imbricate sword-fern, narrow-leaf sword fern, rock sword-fern |
Dudley's sword fern |
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Stems | ascending to erect. |
erect. |
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Leaves | erect to arching back at tip, 2–8 dm; bulblets absent. |
monomorphic, arching, 2–10 dm; bulblets absent. |
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Petiole | 1/4–1/3 length of leaf; scales abruptly diminishing in size distally and falling off early but retaining conspicuous tuft of brown scales at base. |
1/5–1/3 length of leaf, densely scaly; scales light brown, gradually diminishing in size distally. |
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Blade | linear-lanceolate to linear, 1-pinnate, base not or slightly narrowed. |
broadly lanceolate, 2-pinnate, base not narrowed. |
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Pinnae | oblong, slenderly lanceolate, or falcate, usually overlapping, in 1 plane or twisted out of plane of blade, 2–4 cm; base oblique, auricles well developed; margins serrulate-spiny with teeth ascending; apex cuspidate or apiculate with subapical teeth smaller than apical tooth; microscales lanceolate to linear with straight or sharply angular projections, sparse, on abaxial surface only. |
narrowly lanceolate, not overlapping, in 1 plane, 3–13 cm; base oblique, apex acute with subapical and apical teeth same size; microscales filiform, lacking projections, sparse abaxially, but longer than in other Polystichum species, forming loosely tangled network over blade and sori (such network only in this species), sparse adaxially. |
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Indusia | entire to sharply dentate. |
ciliate. |
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Spores | dark brown. |
brown. |
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Pinnules | ± stalked, linear-falcate to oblique-rhombic, acroscopic auricle well developed on proximal pinnules; margins spinulose-dentate; apex acute. |
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2n | = 82. |
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Polystichum imbricans |
Polystichum dudleyi |
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Habitat | Moist forests | |||||
Elevation | 0–100 m (0–300 ft) | |||||
Distribution |
CA; OR; WA; BC; only in the flora
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CA
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Discussion | Polystichum imbricans is one of the postulated ancestors of two allopolyploids, P. californicum and P. scopulinum (D. H. Wagner 1979). Relationships to P. munitum are discussed under that species. Subspecies 2. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Polystichum dudleyi is confined to coastal central California. Hybrids with P. californicum are relatively frequent where these species occur together. These hybrids would key here but, unlike P. dudleyi, they are less divided and have aborted sporangia. The sterile diploid hybrid with P. munitum is also frequent in areas of sympatry. It is indistinguishable from P. californicum except for malformed sporangia and chromosome number (W. H. Wagner Jr. 1973). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 2. | FNA vol. 2. | ||||
Parent taxa | Dryopteridaceae > Polystichum | Dryopteridaceae > Polystichum | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Aspidium munitum var. imbricans, P. munitum subsp. imbricans, P. munitum var. imbricans | P. aculeatum var. dudleyi | ||||
Name authority | (D. C. Eaton) D. H. Wagner: Pteridologia 1: 50. (1979) | Maxon: J. Wash. Acad. Sci. 8: 620. (1918) | ||||
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