The green links below add additional plants to the comparison table. Blue links lead to other Web sites.
enable glossary links

imbricate sword-fern, narrow-leaf sword fern, rock sword-fern

Dudley's sword fern

Stems

ascending to erect.

erect.

Leaves

erect to arching back at tip, 2–8 dm;

bulblets absent.

monomorphic, arching, 2–10 dm;

bulblets absent.

Petiole

1/4–1/3 length of leaf;

scales abruptly diminishing in size distally and falling off early but retaining conspicuous tuft of brown scales at base.

1/5–1/3 length of leaf, densely scaly;

scales light brown, gradually diminishing in size distally.

Blade

linear-lanceolate to linear, 1-pinnate, base not or slightly narrowed.

broadly lanceolate, 2-pinnate, base not narrowed.

Pinnae

oblong, slenderly lanceolate, or falcate, usually overlapping, in 1 plane or twisted out of plane of blade, 2–4 cm;

base oblique, auricles well developed;

margins serrulate-spiny with teeth ascending;

apex cuspidate or apiculate with subapical teeth smaller than apical tooth;

microscales lanceolate to linear with straight or sharply angular projections, sparse, on abaxial surface only.

narrowly lanceolate, not overlapping, in 1 plane, 3–13 cm;

base oblique, apex acute with subapical and apical teeth same size;

microscales filiform, lacking projections, sparse abaxially, but longer than in other Polystichum species, forming loosely tangled network over blade and sori (such network only in this species), sparse adaxially.

Indusia

entire to sharply dentate.

ciliate.

Spores

dark brown.

brown.

Pinnules

± stalked, linear-falcate to oblique-rhombic, acroscopic auricle well developed on proximal pinnules;

margins spinulose-dentate;

apex acute.

2n

= 82.

Polystichum imbricans

Polystichum dudleyi

Habitat Moist forests
Elevation 0–100 m (0–300 ft)
Distribution
from FNA
CA; OR; WA; BC; only in the flora
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
Discussion

Polystichum imbricans is one of the postulated ancestors of two allopolyploids, P. californicum and P. scopulinum (D. H. Wagner 1979). Relationships to P. munitum are discussed under that species.

Subspecies 2.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Polystichum dudleyi is confined to coastal central California. Hybrids with P. californicum are relatively frequent where these species occur together. These hybrids would key here but, unlike P. dudleyi, they are less divided and have aborted sporangia. The sterile diploid hybrid with P. munitum is also frequent in areas of sympatry. It is indistinguishable from P. californicum except for malformed sporangia and chromosome number (W. H. Wagner Jr. 1973).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Pinnae oblong, less than 5 times longer than wide, adaxial surfaces facing upward and twisted out of plane of blade; leaves stiffly erect to tip; at base of boulders or in cliff crevices in exposed sites; California to British Columbia.
subsp. imbricans
1. Pinnae narrowly lanceolate, more than 5 times longer than wide, generally in 1 plane; leaf tips arching back; on forest floor in shade; only in California.
subsp. curtum
Source FNA vol. 2. FNA vol. 2.
Parent taxa Dryopteridaceae > Polystichum Dryopteridaceae > Polystichum
Sibling taxa
P. acrostichoides, P. aleuticum, P. andersonii, P. braunii, P. californicum, P. dudleyi, P. kruckebergii, P. kwakiutlii, P. lemmonii, P. lonchitis, P. microchlamys, P. munitum, P. scopulinum, P. setigerum
P. acrostichoides, P. aleuticum, P. andersonii, P. braunii, P. californicum, P. imbricans, P. kruckebergii, P. kwakiutlii, P. lemmonii, P. lonchitis, P. microchlamys, P. munitum, P. scopulinum, P. setigerum
Subordinate taxa
P. imbricans subsp. curtum, P. imbricans subsp. imbricans
Synonyms Aspidium munitum var. imbricans, P. munitum subsp. imbricans, P. munitum var. imbricans P. aculeatum var. dudleyi
Name authority (D. C. Eaton) D. H. Wagner: Pteridologia 1: 50. (1979) Maxon: J. Wash. Acad. Sci. 8: 620. (1918)
Web links