Polypodium californicum |
Polypodium amorphum |
|
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California polypody |
irregular polypody, irregular polypody fern, Pacific polypody |
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Stems | dull or inconspicuously glaucous, moderately stout, to 10 mm diam., acrid or bland-tasting; scales uniformly brown or slightly darker near point of attachment, lanceolate-ovate, symmetric, margins entire to erose. |
often whitish pruinose, slender, to 6 mm diam., acrid-tasting; scales weakly bicolored, lanceolate, contorted distally, bases and margins light brown, sometimes with dark central stripe, margins often coarsely dentate. |
Leaves | to 70 cm. |
to 30 cm. |
Petiole | usually slender, to 3 mm diam. |
slender, to 1.5 mm diam. |
Blade | deltate to lanceolate-ovate, pinnatifid, usually widest near base, to 20 cm wide, leathery to herbaceous; rachis sparsely scaly to glabrescent abaxially, puberulent adaxially; scales deltate to ovate, usually more than 10 cells wide. |
oblong to rarely deltate, pinnatifid, usually widest near middle, occasionally at or near base, to 4 cm wide, somewhat leathery; rachis sparsely scaly to glabrescent abaxially, glabrous adaxially; scales lanceolate-ovate, usually more than 6 cells wide. |
Segments | linear-lanceolate to oblong, usually less than 15 mm wide; margins serrate; apex widely obtuse to rarely attenuate; midrib puberulent adaxially. |
oblong, less than 12 mm wide; margins entire to crenulate; apex rounded to broadly acute; midrib glabrous adaxially. |
Sori | midway between margin and midrib or slightly closer to midrib, usually less than 3 mm diam., oval when immature. |
midway between margin and midrib to nearly marginal, less than 3 mm diam., circular when immature. |
Spores | less than 58 µm, verrucose, with surface projections to 3 µm. 2n = 74. |
more than 58 µm, rugose to verrucose, surface projections less than 3 µm tall. |
Venation | weakly to conspicuously anastomosing, most segments containing several areoles, often forming 1 row. |
free. |
Sporangiasters | absent. |
present, usually less than 40 per sorus, heads covered with glandular hairs. |
2n | = 74. |
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Polypodium californicum |
Polypodium amorphum |
|
Phenology | Sporulating early winter–spring. | Sporulating summer–fall. |
Habitat | Cliffs and soil on rocky slopes, on a variety of substrates but usually igneous | Cliffs and rocky slopes, usually on igneous substrates |
Elevation | 0–1500 m. (0–4900 ft.) | 0–1800 m (0–5900 ft) |
Distribution |
CA; Mexico in Baja California
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OR; WA; BC
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Discussion | R. M. Lloyd and F. A. Lang (1964) recognized two cytotypes within Polypodium californicum. The tetraploid has proved to be an allopolyploid involving P. californicum and P. glycyrrhiza and is treated here as a separate species, P. calirhiza, following S. A. Whitmore and A. R. Smith (1991). Polypodium californicum can be confused with P. calirhiza, but it usually can be distinguished by blade shape, venation, spore size, and geographic distribution. D. S. Barrington et al. (1986) reported that spores of northern populations of P. californicum can be as large as those of P. calirhiza, but the former species has veins forming more areoles per segment than does the latter. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The diploid Polypodium amorphum is one of the progenitors of allotetraploid P. hesperium, and these two species are occasionally sympatric. Although P. amorphum can be mistaken for P. hesperium, consistent differences exist for separating these two species (see comments under P. hesperium). Hybridization between P. amorphum and P. hesperium results in triploid individuals with misshapen spores (F. A. Lang 1971). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 2. | FNA vol. 2. |
Parent taxa | Polypodiaceae > Polypodium | Polypodiaceae > Polypodium |
Sibling taxa | ||
Synonyms | P. californicum var. kaulfussii, P. vulgare subsp. californicum, P. vulgare var. kaulfussii | P. montense |
Name authority | Kaulfuss: Enum. Filic. 102. (1824) | Suksdorf: Werdenda 1: 16. (1927) |
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