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Johnston's knotweed, knotweed, sawatch knotweed

Shasta knotweed

Habit Herbs. Subshrubs.
Stems

erect, green to brownish, simple or branched from base, not wiry, 4–50 cm, glabrous or papillose-scabrid-ulous.

prostrate to ascending, brown, branched, gnarled, not wiry, 5–40 cm, glabrous.

Leaves

uniformly distributed, articulated to ocreae, basal leaves persistent or caducous, distal leaves abruptly or gradually reduced to bracts;

ocrea 4–10 mm, glabrous or papillose-scabridulous, proximal part cylindric, distal part lacerate or disintegrating into a few persistent fibers;

petiole 0.1–2 mm;

blade 1-veined, not pleated, linear or narrowly oblong to oblanceolate, 8–45 × 2–8(–12) mm, margins revolute or flat, smooth or papillose-denticulate, apex acute, mucronate, rarely obtuse.

± uniformly distributed, articulated to ocreae, basal leaves caducous or persistent, distal leaves not reduced in size;

ocrea 3–5 mm, glabrous, proximal part cylindric, distal part membranous, deciduous with leaves;

petiole 0–0.5 mm;

blade 1-veined, without pleats, lanceolate to elliptic, 5–25 × 2–5 mm, coriaceous, margins revolute, smooth, apex acute.

Inflorescences

axillary or axillary and terminal, spikelike, elongate;

cymes widely spaced along branches, 2–4-flowered.

axillary;

cymes in distal axils, 2–6-flowered.

Pedicels

enclosed in or exserted from ocreae, erect, 1–4 mm.

enclosed in or slightly exserted from ocreae, erect, 2–4 mm.

Flowers

closed or at least some open;

perianth (2.5–)4 mm;

tube 20–40% of perianth length;

tepals overlapping, greenish or reddish, sometimes flushed with purple, with white or pink margins, sepaloid or petaloid, oblong or oblong-elliptic, cucullate, navicular, apex rounded;

midveins unbranched or with few branches at base;

stamens 3–8.

open or semi-open;

perianth 5–9 mm;

tube 7–15% of perianth length, tepals partially overlapping, uniformly pink or white, petaloid, ovate to ovate-round, apex rounded;

midveins with numerous branched lateral veins;

stamens 8.

Achenes

enclosed in perianth, black, elliptic or ovate, 2.5–3.3 mm, faces subequal, shiny, smooth.

enclosed in or slightly exserted from perianth, brown, narrowly ovate, 3–4 mm, faces subequal, shiny, smooth.

Polygonum sawatchense

Polygonum shastense

Phenology Flowering Jul–Sep.
Habitat Rocky or gravelly slopes
Elevation 2100-3400 m (6900-11200 ft)
Distribution
from FNA
AZ; CA; CO; ID; MT; ND; NE; NM; NV; OR; SD; UT; WA; WY; AB; BC; SK
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; NV; OR
[WildflowerSearch map]
[BONAP county map]
Discussion

Subspecies 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Polygonum shastense may be cultivated in rock gardens in rock crevices with favorable water regime and shaded in summer.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Stems and ocreae glabrous; leaf blades linear-lanceolate to narrowly oblong or oblanceolate, margins smooth; flowers closed
subsp. sawatchense
1. Stems and ocreae papillose-scabridulous; leaf blades elliptic or oblong-elliptic, margins papillose-denticulate; at least some flowers open
subsp. oblivium
Source FNA vol. 5, p. 568. FNA vol. 5, p. 563.
Parent taxa Polygonaceae > subfam. Polygonoideae > Polygonum > sect. Duravia Polygonaceae > subfam. Polygonoideae > Polygonum > sect. Duravia
Sibling taxa
P. achoreum, P. argyrocoleon, P. austiniae, P. aviculare, P. bidwelliae, P. bolanderi, P. californicum, P. cascadense, P. douglasii, P. engelmannii, P. erectum, P. fowleri, P. glaucum, P. heterosepalum, P. hickmanii, P. humifusum, P. majus, P. marinense, P. minimum, P. nuttallii, P. oxyspermum, P. paronychia, P. parryi, P. patulum, P. plebeium, P. polygaloides, P. ramosissimum, P. shastense, P. spergulariiforme, P. striatulum, P. tenue, P. utahense
P. achoreum, P. argyrocoleon, P. austiniae, P. aviculare, P. bidwelliae, P. bolanderi, P. californicum, P. cascadense, P. douglasii, P. engelmannii, P. erectum, P. fowleri, P. glaucum, P. heterosepalum, P. hickmanii, P. humifusum, P. majus, P. marinense, P. minimum, P. nuttallii, P. oxyspermum, P. paronychia, P. parryi, P. patulum, P. plebeium, P. polygaloides, P. ramosissimum, P. sawatchense, P. spergulariiforme, P. striatulum, P. tenue, P. utahense
Subordinate taxa
P. sawatchense subsp. oblivium, P. sawatchense subsp. sawatchense
Name authority Small: Bull. Torrey Bot. Club 20: 213, plate 156. (1893) W. H. Brewer: Proc. Amer. Acad. Arts 8: 400. (1872)
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