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bushy knotweed, yellow knotweed, yellow-flower knotweed

close-flower knotweed, close-flower knotweed (ssp. confertiflorum), Kellogg's knotweed (ssp. kelloggii), milkwort knotweed, polygala knotweed, whitemargin knotweed

Habit Plants yellowish green or bluish green, (dark brown to black after drying in subsp. prolificum), heterophyllous or homophyllous. Herbs.
Stems

erect, usually profusely branched in distal 1/2, not wiry, 10–100(–200) cm.

erect, green, usually divaricately branched, rarely simple, ± wiry, (2–)6–20(–25) cm, glabrous.

Leaves

ocrea 6–12(–15) mm, proximal part cylindric, distal part silvery, soon disintegrating into persistent brown fibers;

petiole 2–4 mm;

blade variable, light yellowish green to bluish green, proximal often caducous, narrowly elliptic, lanceolate, or oblanceolate, rarely ovate, 8–70 × 4–18(–35) mm, margins flat, apex acute to acuminate or obtuse;

distal leaves, either overtopping or shorter than or equaling flowers.

uniformly distributed, articulated to ocreae, basal leaves often caducous, distal leaves abruptly reduced to bracts;

ocrea 4–8 mm, glabrous, proximal part cylindric, distal part silvery, with inconspicuous veins, lacerate;

petiole absent;

blade 3-veined, lateral veins sometimes inconspicuous, without pleats, narrowly linear, 10–40 × 1–2.5 mm, margins ± revolute, smooth, apex acute or mucronate.

Inflorescences

axillary or axillary and terminal, spikelike;

cymes uniformly distributed or crowded toward tips of branches, 2–5-flowered.

axillary and terminal, spikelike, subglobose to cylindric;

cymes in most axils or crowded distally, 1–3-flowered.

Pedicels

enclosed in or exserted from ocreae, 1–6 mm.

enclosed in ocreae, erect, 0.1–2 mm, sometimes absent.

Flowers

closed;

perianth (2–)2.2–3.6(–4) mm;

tube 20–38% of perianth length;

tepals overlapping, greenish yellow with greenish yellow or yellow, rarely pink or white, margins, petaloid or sepaloid, not keeled, elliptic to oblong, cucullate;

midveins thickened or not;

stamens 3–6(–8).

mostly closed;

perianth 1.5–3 mm;

tube 19–40% of perianth length;

tepals overlapping, uniformly white, pink, or red, petaloid, oblong-lanceolate, ± navicular, apex acute to acuminate;

midveins usually unbranched or with 2 lateral branches proximally, moderately to strongly thickened, tepals appearing ± keeled;

stamens 3–8.

Achenes

enclosed in or exserted from perianth, dark brown, ovate, 3-gonous, 1.6–3.5 mm, faces subequal, concave, apex not beaked, edges straight, shiny or dull, usually smooth to roughened, sometimes uniformly or obscurely tubercled; late-season achenes common, 4–15 mm.

enclosed in perianth, light yellow, light brown, or greenish brown to dark brown, ovate to lanceolate, 1.3–2.5 mm, faces subequal, shiny or dull, smooth or reticulate with longitudinal ridges.

Polygonum ramosissimum

Polygonum polygaloides

Distribution
from FNA
AR; AZ; CA; CO; CT; DE; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NS; NT; ON; PE; QC; SK
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; CO; ID; MT; NM; NV; OR; UT; WA; WY; AB; BC; SK
[WildflowerSearch map]
[BONAP county map]
Discussion

Subspecies 2 (2 in the flora).

Polygonum ramosissimum exhibits considerable morphological complexity and is similar in difficulty to the P. aviculare complex. Further research is necessary to understand the infraspecific variability of this species (M. Costea and F. J. Tardif 2003b).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 4 (4 in the flora).

J. C. Hickman’s (1993c) treatment of the Polygonum polygaloides complex is provisionally accepted here. Most of the intermediate specimens occur between subspp. confertiflorum, esotericum, and kelloggii. Alternatively, P. polygaloides could be recognized in the narrow sense and the three other taxa could be treated as subspecies of a separate P. kelloggii, the earliest available binomial.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Plants heterophyllous, usually yellowish green when fresh or dried; apices of leaf blades acute to acuminate; pedicels 2.5-6 mm
subsp. ramosissimum
1. Plants homophyllous, bluish green when fresh, dark brown or black after drying; apices of leaf blades rounded or obtuse; pedicels 1-2 mm
subsp. prolificum
1. Margins of bracts green, if white then scarious border less than 0.2 mm wide
→ 2
1. Margins of bracts white, scarious border (0.2-)0.25-1 mm wide
→ 3
2. Achenes lanceolate, 2-2.5 mm; bracts elliptic-lanceolate, appressed, rigid; stamens 5-8
subsp. esotericum
2. Achenes ovate, 1.3-1.7 mm; bracts linear to linear-lanceolate, ± patent, soft; stamens 3
subsp. kelloggii
3. Inflorescences narrowly cylindric, continuous from bases of branches or stems; achenes 2-2.5 mm, lanceolate
subsp. esotericum
3. Inflorescences round to ovate or cylindric, confined to tips of branches, rarely continuous from bases of branches or stems; achenes 1.3-2.1 mm, ovate-lanceolate to ovate
→ 4
4. Inflorescences round to ovate; scarious borders of bracts 0.5-1 mm wide; stamens8
subsp. polygaloides
4. Inflorescences ovate to cylindric; scarious borders of bracts 0.25-0.4 mm wide; stamens 3
subsp. confertiflorum
Source FNA vol. 5, p. 551. FNA vol. 5, p. 565.
Parent taxa Polygonaceae > subfam. Polygonoideae > Polygonum > sect. Polygonum Polygonaceae > subfam. Polygonoideae > Polygonum > sect. Duravia
Sibling taxa
P. achoreum, P. argyrocoleon, P. austiniae, P. aviculare, P. bidwelliae, P. bolanderi, P. californicum, P. cascadense, P. douglasii, P. engelmannii, P. erectum, P. fowleri, P. glaucum, P. heterosepalum, P. hickmanii, P. humifusum, P. majus, P. marinense, P. minimum, P. nuttallii, P. oxyspermum, P. paronychia, P. parryi, P. patulum, P. plebeium, P. polygaloides, P. sawatchense, P. shastense, P. spergulariiforme, P. striatulum, P. tenue, P. utahense
P. achoreum, P. argyrocoleon, P. austiniae, P. aviculare, P. bidwelliae, P. bolanderi, P. californicum, P. cascadense, P. douglasii, P. engelmannii, P. erectum, P. fowleri, P. glaucum, P. heterosepalum, P. hickmanii, P. humifusum, P. majus, P. marinense, P. minimum, P. nuttallii, P. oxyspermum, P. paronychia, P. parryi, P. patulum, P. plebeium, P. ramosissimum, P. sawatchense, P. shastense, P. spergulariiforme, P. striatulum, P. tenue, P. utahense
Subordinate taxa
P. ramosissimum subsp. prolificum, P. ramosissimum subsp. ramosissimum
P. polygaloides subsp. confertiflorum, P. polygaloides subsp. esotericum, P. polygaloides subsp. kelloggii, P. polygaloides subsp. polygaloides
Name authority Michaux: Fl. Bor.-Amer. 1: 237. (1803) Meisner: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 14: 101. (1856)
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