Polygonum polygaloides |
Polygonum douglasii |
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close-flower knotweed, close-flower knotweed (ssp. confertiflorum), Kellogg's knotweed (ssp. kelloggii), milkwort knotweed, polygala knotweed, whitemargin knotweed |
Douglas' knotweed, renouée de Douglas |
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Habit | Herbs. | Herbs. | ||||||||||||||||
Stems | erect, green, usually divaricately branched, rarely simple, ± wiry, (2–)6–20(–25) cm, glabrous. |
erect, green, simple or branched, not wiry, 5–80 cm, glabrous or sparsely papillose-scabridulous. |
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Leaves | uniformly distributed, articulated to ocreae, basal leaves often caducous, distal leaves abruptly reduced to bracts; ocrea 4–8 mm, glabrous, proximal part cylindric, distal part silvery, with inconspicuous veins, lacerate; petiole absent; blade 3-veined, lateral veins sometimes inconspicuous, without pleats, narrowly linear, 10–40 × 1–2.5 mm, margins ± revolute, smooth, apex acute or mucronate. |
uniformly distributed, articulated to ocreae, basal leaves caducous, distal leaves abruptly reduced to bracts; ocrea 6–12 mm, glabrous or minutely papillose-scabridulous, proximal part cylindric, distal part hyaline, lacerate; petiole 0.1–2 mm; blade 1-veined, not pleated, linear, narrow-oblong, or oblanceolate, 15–55 × 2–8(–12) mm, margins revolute, smooth or papillose-denticulate; apex acute to mucronate. |
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Inflorescences | axillary and terminal, spikelike, subglobose to cylindric; cymes in most axils or crowded distally, 1–3-flowered. |
axillary and terminal, spikelike, elongate; cymes widely spaced along branches, 2–4-flowered. |
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Pedicels | enclosed in ocreae, erect, 0.1–2 mm, sometimes absent. |
mostly exserted from ocreae, reflexed, 2–6 mm. |
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Flowers | mostly closed; perianth 1.5–3 mm; tube 19–40% of perianth length; tepals overlapping, uniformly white, pink, or red, petaloid, oblong-lanceolate, ± navicular, apex acute to acuminate; midveins usually unbranched or with 2 lateral branches proximally, moderately to strongly thickened, tepals appearing ± keeled; stamens 3–8. |
closed; perianth 3–4.5 mm; tube 20–28% of perianth length; tepals overlapping, green to tannish with white or pink margins, petaloid, oblong, cucullate, navicular, apex rounded; midveins usually branched, rarely unbranched; stamens 8. |
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Achenes | enclosed in perianth, light yellow, light brown, or greenish brown to dark brown, ovate to lanceolate, 1.3–2.5 mm, faces subequal, shiny or dull, smooth or reticulate with longitudinal ridges. |
enclosed in perianth, black, elliptic or oblong to ovate, 3–4(–4.5) mm, faces subequal, shiny or dull, smooth or minutely striate-tubercled. |
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Polygonum polygaloides |
Polygonum douglasii |
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Phenology | Flowering Jun–Oct. | |||||||||||||||||
Habitat | Dry, often disturbed places, rock outcrops, sandy ground | |||||||||||||||||
Elevation | 300-3000 m (1000-9800 ft) | |||||||||||||||||
Distribution |
AZ; CA; CO; ID; MT; NM; NV; OR; UT; WA; WY; AB; BC; SK
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AZ; CA; CO; IA; ID; MI; MN; MT; NE; NH; NM; NV; NY; OR; SD; UT; VA; VT; WA; WY; AB; BC; MB; ON; QC; SK; YT
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Discussion | Subspecies 4 (4 in the flora). J. C. Hickman’s (1993c) treatment of the Polygonum polygaloides complex is provisionally accepted here. Most of the intermediate specimens occur between subspp. confertiflorum, esotericum, and kelloggii. Alternatively, P. polygaloides could be recognized in the narrow sense and the three other taxa could be treated as subspecies of a separate P. kelloggii, the earliest available binomial. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Five taxa that have been included in Polygonum douglasii (E. Murray 1982; J. C. Hickman 1984; J. T. Kartesz and K. N. Gandhi 1990) are treated here as distinct species: P. austiniae, P. majus, P. nuttallii, P. sawatchense, and P. spergulariiforme. Hickman noted extensive intergradation and numerous intermediate specimens among those sympatric elements, but qualitative or quantitative characters allow reliable discrimination in most cases (M. Costea and F. J. Tardif 2005), and species are here circumscribed similar to C. L. Hitchcock (1964). Greene described var. latifolium as having leaf blades and achenes broader than those of var. douglasii. C. L. Hitchcock (1964) recognized the former, but the characters used to distinguish it appear to vary continuously, and reliable separation is not possible. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 5, p. 565. | FNA vol. 5, p. 567. | ||||||||||||||||
Parent taxa | Polygonaceae > subfam. Polygonoideae > Polygonum > sect. Duravia | Polygonaceae > subfam. Polygonoideae > Polygonum > sect. Duravia | ||||||||||||||||
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Subordinate taxa | ||||||||||||||||||
Synonyms | P. douglasii var. latifolium, P. emaciatum, P. montanum, P. tenue var. commune, P. tenue var. latifolium | |||||||||||||||||
Name authority | Meisner: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 14: 101. (1856) | Greene: Bull. Calif. Acad. Sci. 1: 125. (1885) | ||||||||||||||||
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