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beach knotweed, beach or black or dune knotweed, black knotweed, dune knotweed

Nuttall's knotweed

Habit Shrubs or subshrubs. Herbs.
Stems

prostrate or ascending, brown, branched, rooting at nodes, not wiry, 10–100 cm, glabrous, covered with remains of lacerate, hyaline ocreae.

spreading to erect, sometimes zigzagged, purplish, simple or branched, wiry, (5–)10–35 cm, papillose-scabridulous.

Leaves

crowded at branch tips, articulated to ocreae, basal leaves caducous or persistent, distal leaves not reduced in size;

ocrea 15–20 mm, glabrous, proximal part cylindric to funnelform, distal part silvery, entire or slightly lacerate, disintegrating into persistent white-gray curly fibers;

petiole 0–0.5 mm;

blade 1-veined, without pleats, linear to oblanceolate, (5–)10–20(–33) × 3–8 mm, coriaceous, margins revolute, smooth, apex acute or mucronate.

evenly distributed or crowded at branch tips, articulated to ocreae, basal leaves persistent, distal leaves gradually reduced to bracts;

ocrea 3–4 mm, papillose-scabridulous, proximal part funnelform, distal part finely lacerate;

petiole 0.1–2 mm;

blade 1-veined, not pleated, linear to narrowly oblong-elliptic, 8–30 × 1–4(–7) mm, margins narrowly revolute, never touching along midrib, smooth, apex acute, mucronate, ± glaucous adaxially.

Inflorescences

axillary;

cymes crowded in distal axils, 2–5-flowered.

axillary and terminal, spikelike, dense;

cymes mostly congested toward tips of branches, 2–3-flowered.

Pedicels

enclosed in ocreae, erect to spreading, 2–5 mm.

enclosed in ocreae, erect to spreading, 2–3 mm.

Flowers

semi-open or open;

perianth (4.5–)6–10 mm;

tube 22–48% of perianth length;

tepals partially overlapping, uniformly pink or white, reddish brown when dried, petaloid, oblong-ovate to ± lanceolate, apex rounded;

midveins pinnately branched;

stamens 8.

semi-open or closed;

perianth 1.8–2.4 mm;

tube 25–33% of perianth length;

tepals overlapping, greenish, white, or pink with pink margins, petaloid, oblong, cucullate, navicular in distal 1/4, apex rounded;

midveins unbranched;

stamens 8.

Achenes

enclosed in or slightly exserted from perianth, black, ovate, 4–5 mm, faces subequal, shiny, smooth.

enclosed in perianth, black, elliptic to ovate, 1.8–2.3 mm, faces subequal, shiny, smooth.

Polygonum paronychia

Polygonum nuttallii

Phenology Flowering Mar–Sep. Flowering May–Oct.
Habitat Coastal sands, scrub along coast Dry prairies, open knolls in lower mountains, open sites in lowland and montane zones, sandy soil
Elevation 0-50 m (0-200 ft) 800-1100 m (2600-3600 ft)
Distribution
from FNA
CA; OR; WA; BC
[WildflowerSearch map]
[BONAP county map]
from FNA
OR; WA; BC
[WildflowerSearch map]
[BONAP county map]
Discussion

Polygonum paronychia may be cultivated in rock gardens in open sites with sandy soil.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

C. L. Hitchcock (1964) suggested that Polygonum nuttallii is but a small-flowered form of P. spergulariiforme. Although morphologically similar, P. nuttallii differs from P. spergulariiforme in some respects, including its wiry, purplish stems, short and funnelform ocreae, adaxially glaucous leaves, longer bracts, shorter fruiting perianth, and achenes.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 5, p. 562. FNA vol. 5, p. 570.
Parent taxa Polygonaceae > subfam. Polygonoideae > Polygonum > sect. Duravia Polygonaceae > subfam. Polygonoideae > Polygonum > sect. Duravia
Sibling taxa
P. achoreum, P. argyrocoleon, P. austiniae, P. aviculare, P. bidwelliae, P. bolanderi, P. californicum, P. cascadense, P. douglasii, P. engelmannii, P. erectum, P. fowleri, P. glaucum, P. heterosepalum, P. hickmanii, P. humifusum, P. majus, P. marinense, P. minimum, P. nuttallii, P. oxyspermum, P. parryi, P. patulum, P. plebeium, P. polygaloides, P. ramosissimum, P. sawatchense, P. shastense, P. spergulariiforme, P. striatulum, P. tenue, P. utahense
P. achoreum, P. argyrocoleon, P. austiniae, P. aviculare, P. bidwelliae, P. bolanderi, P. californicum, P. cascadense, P. douglasii, P. engelmannii, P. erectum, P. fowleri, P. glaucum, P. heterosepalum, P. hickmanii, P. humifusum, P. majus, P. marinense, P. minimum, P. oxyspermum, P. paronychia, P. parryi, P. patulum, P. plebeium, P. polygaloides, P. ramosissimum, P. sawatchense, P. shastense, P. spergulariiforme, P. striatulum, P. tenue, P. utahense
Synonyms P. intermedium, P. douglasii subsp. nuttallii
Name authority Chamisso & Schlechtendal: Linnaea 3: 51. (1828) Small: Mongr. Amer. Sp. Polygonum, 132, plate 53. (1895)
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