Polygonum minimum |
Polygonum aviculare |
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broad-leaf knotweed, broad-leaf or leafy dwarf knotweed, leafy dwarf knotweed, little mountain knotweed, zigzag knotweed |
birdweed, common knotgrass, common knotweed, doorweed, dooryard knotweed, knotgrass, knotweed, lowgrass, pigweed, prostrate knotweed, renouée des oiseaux, yard knotweed |
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Habit | Herbs. | Plants green or bluish green, green after drying, sometimes whitish from powdery mildew, homophyllous or heterophyllous. | ||||||||||||||||||||||||||||
Stems | prostrate to erect, often zigzagged, reddish brown, simple or branched from base, wiry, 2–30 cm, papillose-scabrid-ulous. |
prostrate to erect, branched, flexuous, 5–200 cm. |
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Leaves | evenly distributed or crowded at branch tips, articulated to ocreae, basal leaves persistent, hardly reduced distally; ocrea 1–4 mm, papillose-scabridulous, proximal part cylindric, distal part entire or dentate-lacerate; petiole 0.1–3 mm; blade 1-veined, not pleated, narrowly elliptic, elliptic, ovate, obovate, or subround, 6–27 × 3–8 mm, margins flat, smooth, irregularly thickened or papillose-denticulate, apex apiculate, green adaxially. |
ocrea 3–15 mm, proximal part cylindric or ± funnelform, distal part silvery, hyaline, soon disintegrating into persistent fibers or nearly completely deciduous; petiole 0.3–9 mm; blade green to gray-green, narrowly elliptic, lanceolate, elliptic, obovate, or spatulate, 6–50(–60) × 0.5–22 mm, margins flat, apex acute, obtuse, or rounded; stem leaves 1–4 times as long as adjacent branch leaves; distal leaves overtopping flowers. |
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Inflorescences | axillary; cymes from near stem and branch bases, sometimes also crowded at branch apices, 1–3-flowered. |
axillary; cymes uniformly distributed or aggregated at tips of stems and branches, 1–6(–8)-flowered. |
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Pedicels | enclosed in ocreae, erect to spreading, 2–3 mm. |
enclosed in or exserted from ocreae, 1.5–5 mm. |
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Flowers | semi-open or closed; perianth 1.8–2.5 mm; tube 22–29% of perianth length; tepals overlapping, greenish with narrow white or pink margins, almost sepaloid, oblong, cucullate, ± navicular, apex rounded; midveins thickened, unbranched; stamens 8. |
closed or semi-open; perianth 1.8–5.5 mm; tube 20–57% of perianth length; tepals overlapping or not, green or reddish brown with white, pink, or red margins, petaloid, not keeled, oblong to obovate, often cucullate in fruit; midveins branched or unbranched, thickened or not; stamens 5–8. |
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Achenes | enclosed in perianth or tip exserted, black, elliptic to ovate, 1.8–2.3 mm, faces subequal, smooth, shiny. |
enclosed in or exserted from perianth, light to dark brown, ovate, (2–)3-gonous, 1.2–4.2 mm, faces subequal or unequal, apex not beaked, edges slightly concave, dull, usually coarsely striate-tubercled, sometimes obscurely tubercled; late-season achenes common or not, 2–5 mm. |
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Polygonum minimum |
Polygonum aviculare |
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Phenology | Flowering Jul–Sep. | |||||||||||||||||||||||||||||
Habitat | Alpine to subalpine sites, open or semibarren soil | |||||||||||||||||||||||||||||
Elevation | 1500-3300 m (4900-10800 ft) | |||||||||||||||||||||||||||||
Distribution |
AK; CA; CO; ID; MT; NV; OR; UT; WA; WY; AB; BC
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AK; AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NL; NS; NT; ON; PE; QC; SK; YT; nearly worldwide
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Discussion | Subspecies 7+ (6 in the flora). Polygonum aviculare is a taxonomically controversial polyploid complex of selfing annuals. Although members of the complex have been considered inbreeders, they possess some structures that make cross pollination possible. Cleistogamous and chasmogamous flowers, heterostyly, protandry, and the capacity to secrete nectar suggest an ancestral mixed-mating system. Isoenzyme studies showed that the complex has an allopolyploid origin (P. Meerts et al. 1998) and has evolved as a swarm of inbreeding lines (“Jordanons”) (J. Gasquez et al. 1978). The six subspecies included here have been treated variously (T. Karlsson 2000; M. Costea and F. J. Tardif 2003). Complex intergradation patterns among them make their recognition at the species level impractical. Multivariate analysis and isoenzyme studies show that populations with intermediate characteristics may occur (Meerts et al. 1990, 1998). Except for subsp. boreale, which occurs in Greenland and Labrador, all subspecies are partially sympatric and their distributions have been influenced greatly by humans. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 5, p. 569. | FNA vol. 5, p. 556. | ||||||||||||||||||||||||||||
Parent taxa | Polygonaceae > subfam. Polygonoideae > Polygonum > sect. Duravia | Polygonaceae > subfam. Polygonoideae > Polygonum > sect. Polygonum | ||||||||||||||||||||||||||||
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Synonyms | P. torreyi | |||||||||||||||||||||||||||||
Name authority | S. Watson: Botany (Fortieth Parallel), 315. (1871) | Linnaeus: Sp. Pl. 1: 362. (1753) | ||||||||||||||||||||||||||||
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