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large Douglas' knotweed, large knotweed, Palouse knotweed, wiry knotweed

knotweed, sharp-fruit knotweed

Habit Herbs. Plants green to blue-green, homophyllous.
Stems

erect, green, simple or branched, ± wiry, 15–60 cm, usually papillose-scabridulous.

prostrate to ascending, branched at proximal and middle nodes, not wiry, 20–100 cm.

Leaves

uniformly distributed, articulated to ocreae, basal leaves often caducous, distal leaves abruptly reduced to bracts, articulated to ocreae;

ocrea 5–12 mm, glabrous or papillose-scabridulous, proximal part cylindric, distal part lacerate or disintegrating into fibers;

petiole 0.1–2 mm;

blade 1-veined, not pleated, linear to narrowly oblong or lanceolate, 15–70 × 2–8 mm, margins revolute, papillose-denticulate, apex acute or mucronate.

ocrea 6.5–12 mm, proximal part cylindric, pruinose, distal part hyaline, soon disintegrating into brown fibers or nearly completely deciduous;

petiole 0–1 mm;

blade green to bluish green, elliptic-lanceolate to linear, 10–35 × 1.5–15 mm, margins flat or narrowly revolute, apex acute;

middle stem leaves slightly larger than adjacent branch leaves, distal leaves overtopping flowers.

Inflorescences

axillary and terminal, spikelike, elongate;

cymes spaced along branches, 2–5-flowered.

axillary, cymes uniformly distributed, 2–7-flowered.

Pedicels

exserted from ocreae, reflexed, 0.5–1 mm.

exserted from ocreae, 2.5–5(–7) mm.

Flowers

open or semi-open;

perianth (3.5–)4–5 mm;

tube 9–17% of perianth length;

tepals overlapping, uniformly white to pink, petaloid, oblong to oblong-obovate, cucullate, navicular in distal 1/4, apex rounded;

midveins unbranched or with short lateral branches;

stamens 8.

semi-open;

perianth 3.5–5.5 mm;

tube 28–39% of perianth length;

tepals slightly overlapping, green, margins white to pink, petaloid, not keeled, oblong to obovate, not cucullate;

veins branched;

stamens 8.

Achenes

enclosed in perianth, black, elliptic, 3.5–5 mm, faces subequal, shiny or dull, smooth or striate-tubercled.

exserted from perianth, pale brown to dark brown, ovate, 3-gonous, (3.5–)4.1–5.5(–6.5) mm, faces subequal, apex not beaked, edges straight, shiny, smooth or with fine tubercles especially toward apex; late-season achenes unknown.

Polygonum majus

Polygonum oxyspermum

Phenology Flowering May–Aug.
Habitat Dry plains, meadows, sometimes on serpentine
Elevation 500-2000 m (1600-6600 ft)
Distribution
from FNA
CA; CO; ID; NV; OR; UT; WA; WY; BC
[WildflowerSearch map]
[BONAP county map]
from FNA
ME; NB; NF; NS; PE; QC; Europe
[BONAP county map]
Discussion

Subspecies 3 (2 in the flora).

The treatment by D. A. Webb and A. O. Chater (1963) of the Polygonum oxyspermum complex is followed here. T. Karlsson (2000) accepted P. raii and P. oxyspermum as distinct species.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Ocreae with prominent veins, distal part disintegrating, veins persistent; leaf blades 5-8 times as long as wide; achenes 4.1-5.5(-6.5) mm, exserted 1-2.5 mm from perianth
subsp. oxyspermum
1. Ocreae without prominent veins, distal part disintegrating, nearly completely deciduous; leaf blades 3.5-5 times as long as wide; achene (3.5-)4-5 mm, exserted 0.5-1.3(-1.8) mm from perianth
subsp. raii
Source FNA vol. 5, p. 570. FNA vol. 5, p. 554.
Parent taxa Polygonaceae > subfam. Polygonoideae > Polygonum > sect. Duravia Polygonaceae > subfam. Polygonoideae > Polygonum > sect. Polygonum
Sibling taxa
P. achoreum, P. argyrocoleon, P. austiniae, P. aviculare, P. bidwelliae, P. bolanderi, P. californicum, P. cascadense, P. douglasii, P. engelmannii, P. erectum, P. fowleri, P. glaucum, P. heterosepalum, P. hickmanii, P. humifusum, P. marinense, P. minimum, P. nuttallii, P. oxyspermum, P. paronychia, P. parryi, P. patulum, P. plebeium, P. polygaloides, P. ramosissimum, P. sawatchense, P. shastense, P. spergulariiforme, P. striatulum, P. tenue, P. utahense
P. achoreum, P. argyrocoleon, P. austiniae, P. aviculare, P. bidwelliae, P. bolanderi, P. californicum, P. cascadense, P. douglasii, P. engelmannii, P. erectum, P. fowleri, P. glaucum, P. heterosepalum, P. hickmanii, P. humifusum, P. majus, P. marinense, P. minimum, P. nuttallii, P. paronychia, P. parryi, P. patulum, P. plebeium, P. polygaloides, P. ramosissimum, P. sawatchense, P. shastense, P. spergulariiforme, P. striatulum, P. tenue, P. utahense
Subordinate taxa
P. oxyspermum subsp. oxyspermum, P. oxyspermum subsp. raii
Synonyms P. coarctatum var. majus, P. douglasii subsp. majus
Name authority (Meisner) Piper: Fl. Palouse Reg., 63. (1901) C. A. Meyer & Bunge ex Ledebour: Index Seminum (Dorpat) 1824: 5. (1824)
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