Polygonum douglasii |
Polygonum paronychia |
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Douglas' knotweed, renouée de Douglas |
beach knotweed, beach or black or dune knotweed, black knotweed, dune knotweed |
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Habit | Herbs. | Shrubs or subshrubs. |
Stems | erect, green, simple or branched, not wiry, 5–80 cm, glabrous or sparsely papillose-scabridulous. |
prostrate or ascending, brown, branched, rooting at nodes, not wiry, 10–100 cm, glabrous, covered with remains of lacerate, hyaline ocreae. |
Leaves | uniformly distributed, articulated to ocreae, basal leaves caducous, distal leaves abruptly reduced to bracts; ocrea 6–12 mm, glabrous or minutely papillose-scabridulous, proximal part cylindric, distal part hyaline, lacerate; petiole 0.1–2 mm; blade 1-veined, not pleated, linear, narrow-oblong, or oblanceolate, 15–55 × 2–8(–12) mm, margins revolute, smooth or papillose-denticulate; apex acute to mucronate. |
crowded at branch tips, articulated to ocreae, basal leaves caducous or persistent, distal leaves not reduced in size; ocrea 15–20 mm, glabrous, proximal part cylindric to funnelform, distal part silvery, entire or slightly lacerate, disintegrating into persistent white-gray curly fibers; petiole 0–0.5 mm; blade 1-veined, without pleats, linear to oblanceolate, (5–)10–20(–33) × 3–8 mm, coriaceous, margins revolute, smooth, apex acute or mucronate. |
Inflorescences | axillary and terminal, spikelike, elongate; cymes widely spaced along branches, 2–4-flowered. |
axillary; cymes crowded in distal axils, 2–5-flowered. |
Pedicels | mostly exserted from ocreae, reflexed, 2–6 mm. |
enclosed in ocreae, erect to spreading, 2–5 mm. |
Flowers | closed; perianth 3–4.5 mm; tube 20–28% of perianth length; tepals overlapping, green to tannish with white or pink margins, petaloid, oblong, cucullate, navicular, apex rounded; midveins usually branched, rarely unbranched; stamens 8. |
semi-open or open; perianth (4.5–)6–10 mm; tube 22–48% of perianth length; tepals partially overlapping, uniformly pink or white, reddish brown when dried, petaloid, oblong-ovate to ± lanceolate, apex rounded; midveins pinnately branched; stamens 8. |
Achenes | enclosed in perianth, black, elliptic or oblong to ovate, 3–4(–4.5) mm, faces subequal, shiny or dull, smooth or minutely striate-tubercled. |
enclosed in or slightly exserted from perianth, black, ovate, 4–5 mm, faces subequal, shiny, smooth. |
Polygonum douglasii |
Polygonum paronychia |
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Phenology | Flowering Jun–Oct. | Flowering Mar–Sep. |
Habitat | Dry, often disturbed places, rock outcrops, sandy ground | Coastal sands, scrub along coast |
Elevation | 300-3000 m (1000-9800 ft) | 0-50 m (0-200 ft) |
Distribution |
AZ; CA; CO; IA; ID; MI; MN; MT; NE; NH; NM; NV; NY; OR; SD; UT; VA; VT; WA; WY; AB; BC; MB; ON; QC; SK; YT
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CA; OR; WA; BC
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Discussion | Five taxa that have been included in Polygonum douglasii (E. Murray 1982; J. C. Hickman 1984; J. T. Kartesz and K. N. Gandhi 1990) are treated here as distinct species: P. austiniae, P. majus, P. nuttallii, P. sawatchense, and P. spergulariiforme. Hickman noted extensive intergradation and numerous intermediate specimens among those sympatric elements, but qualitative or quantitative characters allow reliable discrimination in most cases (M. Costea and F. J. Tardif 2005), and species are here circumscribed similar to C. L. Hitchcock (1964). Greene described var. latifolium as having leaf blades and achenes broader than those of var. douglasii. C. L. Hitchcock (1964) recognized the former, but the characters used to distinguish it appear to vary continuously, and reliable separation is not possible. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Polygonum paronychia may be cultivated in rock gardens in open sites with sandy soil. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 5, p. 567. | FNA vol. 5, p. 562. |
Parent taxa | Polygonaceae > subfam. Polygonoideae > Polygonum > sect. Duravia | Polygonaceae > subfam. Polygonoideae > Polygonum > sect. Duravia |
Sibling taxa | ||
Synonyms | P. douglasii var. latifolium, P. emaciatum, P. montanum, P. tenue var. commune, P. tenue var. latifolium | |
Name authority | Greene: Bull. Calif. Acad. Sci. 1: 125. (1885) | Chamisso & Schlechtendal: Linnaea 3: 51. (1828) |
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