Polygonum douglasii |
Polygonum heterosepalum |
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Douglas' knotweed, renouée de Douglas |
dwarf desert knotweed, odd-sepal knotweed |
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Habit | Herbs. | Herbs, compact, often cushion-like. |
Stems | erect, green, simple or branched, not wiry, 5–80 cm, glabrous or sparsely papillose-scabridulous. |
erect, green or reddish, simple or branched near base, not wiry, 1.5–5 cm, glabrous. |
Leaves | uniformly distributed, articulated to ocreae, basal leaves caducous, distal leaves abruptly reduced to bracts; ocrea 6–12 mm, glabrous or minutely papillose-scabridulous, proximal part cylindric, distal part hyaline, lacerate; petiole 0.1–2 mm; blade 1-veined, not pleated, linear, narrow-oblong, or oblanceolate, 15–55 × 2–8(–12) mm, margins revolute, smooth or papillose-denticulate; apex acute to mucronate. |
uniformly distributed, dense, not articulated to ocreae, basal leaves persistent or caducous, distal leaves gradually reduced to bracts; ocrea 3–6 mm, glabrous, proximal part cylindric, distal part disintegrating almost to base, with whitish, straight, rigid fibers; petiole absent; blade 3-veined, without pleats, linear to lanceolate, 10–20 × 0.6–2.7 mm, margins revolute, smooth, apex spine-tipped. |
Inflorescences | axillary and terminal, spikelike, elongate; cymes widely spaced along branches, 2–4-flowered. |
axillary; cymes in most axils, 2–3-flowered. |
Pedicels | mostly exserted from ocreae, reflexed, 2–6 mm. |
enclosed in ocreae, erect, 0.1–1 mm. |
Flowers | closed; perianth 3–4.5 mm; tube 20–28% of perianth length; tepals overlapping, green to tannish with white or pink margins, petaloid, oblong, cucullate, navicular, apex rounded; midveins usually branched, rarely unbranched; stamens 8. |
closed; perianth 2.3–2.7 mm; tube 3–7% of perianth length; tepals overlapping, whitish with whitish or pink margins, petaloid, oblong, navicular, dimorphic, outer 2 shorter than inner 3, outer 2 0.8–1.2 mm, inner 2.3–2.7 mm, papillose at base, apex acute or acuminate; midveins unbranched; stamens 5–6. |
Achenes | enclosed in perianth, black, elliptic or oblong to ovate, 3–4(–4.5) mm, faces subequal, shiny or dull, smooth or minutely striate-tubercled. |
enclosed in perianth, olive brown, narrowly ovate to ovate-lanceolate, 1.5–2 mm, faces subequal, shiny, smooth. |
Polygonum douglasii |
Polygonum heterosepalum |
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Phenology | Flowering Jun–Oct. | Flowering Jun–Aug. |
Habitat | Dry, often disturbed places, rock outcrops, sandy ground | Dry waste ground, open flats in sagebrush plains, ponderosa pine forests |
Elevation | 300-3000 m (1000-9800 ft) | 1000-1500 m (3300-4900 ft) |
Distribution |
AZ; CA; CO; IA; ID; MI; MN; MT; NE; NH; NM; NV; NY; OR; SD; UT; VA; VT; WA; WY; AB; BC; MB; ON; QC; SK; YT
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ID; NV; OR |
Discussion | Five taxa that have been included in Polygonum douglasii (E. Murray 1982; J. C. Hickman 1984; J. T. Kartesz and K. N. Gandhi 1990) are treated here as distinct species: P. austiniae, P. majus, P. nuttallii, P. sawatchense, and P. spergulariiforme. Hickman noted extensive intergradation and numerous intermediate specimens among those sympatric elements, but qualitative or quantitative characters allow reliable discrimination in most cases (M. Costea and F. J. Tardif 2005), and species are here circumscribed similar to C. L. Hitchcock (1964). Greene described var. latifolium as having leaf blades and achenes broader than those of var. douglasii. C. L. Hitchcock (1964) recognized the former, but the characters used to distinguish it appear to vary continuously, and reliable separation is not possible. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 5, p. 567. | FNA vol. 5, p. 563. |
Parent taxa | Polygonaceae > subfam. Polygonoideae > Polygonum > sect. Duravia | Polygonaceae > subfam. Polygonoideae > Polygonum > sect. Duravia |
Sibling taxa | ||
Synonyms | P. douglasii var. latifolium, P. emaciatum, P. montanum, P. tenue var. commune, P. tenue var. latifolium | |
Name authority | Greene: Bull. Calif. Acad. Sci. 1: 125. (1885) | M. Peck & Ownbey: Madroño 10: 250. (1950) |
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