Polygonum aviculare |
Polygonum utahense |
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birdweed, common knotgrass, common knotweed, doorweed, dooryard knotweed, knotgrass, knotweed, lowgrass, pigweed, prostrate knotweed, renouée des oiseaux, yard knotweed |
Utah knotweed |
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Habit | Plants green or bluish green, green after drying, sometimes whitish from powdery mildew, homophyllous or heterophyllous. | Herbs. | ||||||||||||||||||||||||||||
Stems | prostrate to erect, branched, flexuous, 5–200 cm. |
erect, green, simple or with few branches, not wiry, 1.5–3.5 cm, papillose-scabridulous. |
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Leaves | ocrea 3–15 mm, proximal part cylindric or ± funnelform, distal part silvery, hyaline, soon disintegrating into persistent fibers or nearly completely deciduous; petiole 0.3–9 mm; blade green to gray-green, narrowly elliptic, lanceolate, elliptic, obovate, or spatulate, 6–50(–60) × 0.5–22 mm, margins flat, apex acute, obtuse, or rounded; stem leaves 1–4 times as long as adjacent branch leaves; distal leaves overtopping flowers. |
uniformly distributed, articulated to ocreae, basal leaves persistent, distal leaves abruptly reduced to bracts; ocrea 2–3.5 mm, papillose-scabridulous, proximal part funnelform, distal part hyaline, lacerate; petiole absent; blade 1-veined, not pleated, linear-subulate, 6–14 × 1 mm, coriaceous, margins revolute, touching along midrib, apex acute, mucronate, papillose. |
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Inflorescences | axillary; cymes uniformly distributed or aggregated at tips of stems and branches, 1–6(–8)-flowered. |
axillary and terminal, spikelike, dense; cymes overlapping, starting almost from stem base, 2–4-flowered. |
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Pedicels | enclosed in or exserted from ocreae, 1.5–5 mm. |
enclosed in ocreae, erect to spreading, 0.4–1 mm. |
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Flowers | closed or semi-open; perianth 1.8–5.5 mm; tube 20–57% of perianth length; tepals overlapping or not, green or reddish brown with white, pink, or red margins, petaloid, not keeled, oblong to obovate, often cucullate in fruit; midveins branched or unbranched, thickened or not; stamens 5–8. |
open; perianth 2–2.5 mm; tube 20–25% of perianth length; tepals overlapping, uniformly white, petaloid, oblong to widely obovate, cucullate, navicular in distal 1/4, apex rounded; midveins branched; stamens 8. |
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Achenes | enclosed in or exserted from perianth, light to dark brown, ovate, (2–)3-gonous, 1.2–4.2 mm, faces subequal or unequal, apex not beaked, edges slightly concave, dull, usually coarsely striate-tubercled, sometimes obscurely tubercled; late-season achenes common or not, 2–5 mm. |
enclosed in perianth, black, elliptic, 1.5–2 mm, faces subequal, shiny, smooth. |
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Polygonum aviculare |
Polygonum utahense |
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Phenology | Flowering Jul–Sep. | |||||||||||||||||||||||||||||
Habitat | Dry, sandy ravines, rocky Navajo sandstone spur | |||||||||||||||||||||||||||||
Elevation | 2300 m (7500 ft) | |||||||||||||||||||||||||||||
Distribution |
AK; AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NL; NS; NT; ON; PE; QC; SK; YT; nearly worldwide
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UT |
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Discussion | Subspecies 7+ (6 in the flora). Polygonum aviculare is a taxonomically controversial polyploid complex of selfing annuals. Although members of the complex have been considered inbreeders, they possess some structures that make cross pollination possible. Cleistogamous and chasmogamous flowers, heterostyly, protandry, and the capacity to secrete nectar suggest an ancestral mixed-mating system. Isoenzyme studies showed that the complex has an allopolyploid origin (P. Meerts et al. 1998) and has evolved as a swarm of inbreeding lines (“Jordanons”) (J. Gasquez et al. 1978). The six subspecies included here have been treated variously (T. Karlsson 2000; M. Costea and F. J. Tardif 2003). Complex intergradation patterns among them make their recognition at the species level impractical. Multivariate analysis and isoenzyme studies show that populations with intermediate characteristics may occur (Meerts et al. 1990, 1998). Except for subsp. boreale, which occurs in Greenland and Labrador, all subspecies are partially sympatric and their distributions have been influenced greatly by humans. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Polygonum utahense has been treated as a synonym of P. sawatchense by some authors (e.g., S. L. Welsh et al. 1987; J. T. Kartesz and C. A. Meacham 1999). Its perianth morphology suggests a relationship with P. cascadense, from which it differs in its dwarf habit and uniformly papillose leaves with margins revolute, touching along midribs. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 5, p. 556. | FNA vol. 5, p. 571. | ||||||||||||||||||||||||||||
Parent taxa | Polygonaceae > subfam. Polygonoideae > Polygonum > sect. Polygonum | Polygonaceae > subfam. Polygonoideae > Polygonum > sect. Duravia | ||||||||||||||||||||||||||||
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Synonyms | P. douglasii var. utahense | |||||||||||||||||||||||||||||
Name authority | Linnaeus: Sp. Pl. 1: 362. (1753) | Brenckle & Cottam: Bull. Univ. Utah, Biol. Ser. 4(4): 3, plate 1. (1940) | ||||||||||||||||||||||||||||
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