Polygonum aviculare |
Polygonum striatulum |
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birdweed, common knotgrass, common knotweed, doorweed, dooryard knotweed, knotgrass, knotweed, lowgrass, pigweed, prostrate knotweed, renouée des oiseaux, yard knotweed |
striped knotweed, Texas knotweed |
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Habit | Plants green or bluish green, green after drying, sometimes whitish from powdery mildew, homophyllous or heterophyllous. | Plants perennial, light green, heterophyllous; rhizomes brown, 0.7–3 cm diam. | ||||||||||||||||||||||||||||
Stems | prostrate to erect, branched, flexuous, 5–200 cm. |
erect to ascending, sparingly branched in distal 1/2, not wiry, 25–60 cm. |
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Leaves | ocrea 3–15 mm, proximal part cylindric or ± funnelform, distal part silvery, hyaline, soon disintegrating into persistent fibers or nearly completely deciduous; petiole 0.3–9 mm; blade green to gray-green, narrowly elliptic, lanceolate, elliptic, obovate, or spatulate, 6–50(–60) × 0.5–22 mm, margins flat, apex acute, obtuse, or rounded; stem leaves 1–4 times as long as adjacent branch leaves; distal leaves overtopping flowers. |
ocrea 6–12 mm, proximal part cylindric, distal part soon disintegrating into brown fibers, later leaving almost no fibrous remains; petiole 0–2 mm; blade light green, linear-lanceolate to oblanceolate, 8–35 × 2–8 mm, margins flat, apex acute or obtuse; stem leaves 2.1–4 times as long as adjacent branch leaves; distal leaves sharply reduced, not overtopping flowers (shorter than or equaling flowers). |
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Inflorescences | axillary; cymes uniformly distributed or aggregated at tips of stems and branches, 1–6(–8)-flowered. |
axillary and terminal, spikelike; cymes in distal nodes, 2–6-flowered. |
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Pedicels | enclosed in or exserted from ocreae, 1.5–5 mm. |
exserted from ocreae, 2–4 mm. |
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Flowers | closed or semi-open; perianth 1.8–5.5 mm; tube 20–57% of perianth length; tepals overlapping or not, green or reddish brown with white, pink, or red margins, petaloid, not keeled, oblong to obovate, often cucullate in fruit; midveins branched or unbranched, thickened or not; stamens 5–8. |
semi-open; perianth 2–3.5 mm; tube 18–25% of perianth length; tepals overlapping, green with white or sometimes pink margins, petaloid, not keeled, oblong to obovate, cucullate; midveins usually unbranched; stamens 7–8. |
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Achenes | enclosed in or exserted from perianth, light to dark brown, ovate, (2–)3-gonous, 1.2–4.2 mm, faces subequal or unequal, apex not beaked, edges slightly concave, dull, usually coarsely striate-tubercled, sometimes obscurely tubercled; late-season achenes common or not, 2–5 mm. |
enclosed in perianth, brown, ovate, 3-gonous, (1.6–)1.8–2.6(–3) mm, faces subequal or unequal, apex not beaked, edges concave, shiny, smooth or roughened; late-season achenes common, 4–6 mm. |
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Polygonum aviculare |
Polygonum striatulum |
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Phenology | Flowering Dec–Mar or Jun–Oct. | |||||||||||||||||||||||||||||
Habitat | Seasonal moist places, sterile prairies, granitic soils | |||||||||||||||||||||||||||||
Elevation | 100-700 m (300-2300 ft) | |||||||||||||||||||||||||||||
Distribution |
AK; AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NL; NS; NT; ON; PE; QC; SK; YT; nearly worldwide
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TX |
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Discussion | Subspecies 7+ (6 in the flora). Polygonum aviculare is a taxonomically controversial polyploid complex of selfing annuals. Although members of the complex have been considered inbreeders, they possess some structures that make cross pollination possible. Cleistogamous and chasmogamous flowers, heterostyly, protandry, and the capacity to secrete nectar suggest an ancestral mixed-mating system. Isoenzyme studies showed that the complex has an allopolyploid origin (P. Meerts et al. 1998) and has evolved as a swarm of inbreeding lines (“Jordanons”) (J. Gasquez et al. 1978). The six subspecies included here have been treated variously (T. Karlsson 2000; M. Costea and F. J. Tardif 2003). Complex intergradation patterns among them make their recognition at the species level impractical. Multivariate analysis and isoenzyme studies show that populations with intermediate characteristics may occur (Meerts et al. 1990, 1998). Except for subsp. boreale, which occurs in Greenland and Labrador, all subspecies are partially sympatric and their distributions have been influenced greatly by humans. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Plants with leaves less-conspicuously veined, flowering August to October, and growing in seasonally moist habitats in central and western Texas are recognized by some authors as Polygonum texense or P. striatulum var. texense (M. Costea and F. J. Tardif 2003). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 5, p. 556. | FNA vol. 5, p. 550. | ||||||||||||||||||||||||||||
Parent taxa | Polygonaceae > subfam. Polygonoideae > Polygonum > sect. Polygonum | Polygonaceae > subfam. Polygonoideae > Polygonum > sect. Polygonum | ||||||||||||||||||||||||||||
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Synonyms | P. striatulum var. texense, P. texense | |||||||||||||||||||||||||||||
Name authority | Linnaeus: Sp. Pl. 1: 362. (1753) | B. L. Robinson: Proc. Boston Soc. Nat. Hist. 31: 263. (1904) | ||||||||||||||||||||||||||||
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