Polygonum aviculare |
Polygonum shastense |
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birdweed, common knotgrass, common knotweed, doorweed, dooryard knotweed, knotgrass, knotweed, lowgrass, pigweed, prostrate knotweed, renouée des oiseaux, yard knotweed |
Shasta knotweed |
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Habit | Plants green or bluish green, green after drying, sometimes whitish from powdery mildew, homophyllous or heterophyllous. | Subshrubs. | ||||||||||||||||||||||||||||
Stems | prostrate to erect, branched, flexuous, 5–200 cm. |
prostrate to ascending, brown, branched, gnarled, not wiry, 5–40 cm, glabrous. |
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Leaves | ocrea 3–15 mm, proximal part cylindric or ± funnelform, distal part silvery, hyaline, soon disintegrating into persistent fibers or nearly completely deciduous; petiole 0.3–9 mm; blade green to gray-green, narrowly elliptic, lanceolate, elliptic, obovate, or spatulate, 6–50(–60) × 0.5–22 mm, margins flat, apex acute, obtuse, or rounded; stem leaves 1–4 times as long as adjacent branch leaves; distal leaves overtopping flowers. |
± uniformly distributed, articulated to ocreae, basal leaves caducous or persistent, distal leaves not reduced in size; ocrea 3–5 mm, glabrous, proximal part cylindric, distal part membranous, deciduous with leaves; petiole 0–0.5 mm; blade 1-veined, without pleats, lanceolate to elliptic, 5–25 × 2–5 mm, coriaceous, margins revolute, smooth, apex acute. |
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Inflorescences | axillary; cymes uniformly distributed or aggregated at tips of stems and branches, 1–6(–8)-flowered. |
axillary; cymes in distal axils, 2–6-flowered. |
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Pedicels | enclosed in or exserted from ocreae, 1.5–5 mm. |
enclosed in or slightly exserted from ocreae, erect, 2–4 mm. |
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Flowers | closed or semi-open; perianth 1.8–5.5 mm; tube 20–57% of perianth length; tepals overlapping or not, green or reddish brown with white, pink, or red margins, petaloid, not keeled, oblong to obovate, often cucullate in fruit; midveins branched or unbranched, thickened or not; stamens 5–8. |
open or semi-open; perianth 5–9 mm; tube 7–15% of perianth length, tepals partially overlapping, uniformly pink or white, petaloid, ovate to ovate-round, apex rounded; midveins with numerous branched lateral veins; stamens 8. |
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Achenes | enclosed in or exserted from perianth, light to dark brown, ovate, (2–)3-gonous, 1.2–4.2 mm, faces subequal or unequal, apex not beaked, edges slightly concave, dull, usually coarsely striate-tubercled, sometimes obscurely tubercled; late-season achenes common or not, 2–5 mm. |
enclosed in or slightly exserted from perianth, brown, narrowly ovate, 3–4 mm, faces subequal, shiny, smooth. |
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Polygonum aviculare |
Polygonum shastense |
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Phenology | Flowering Jul–Sep. | |||||||||||||||||||||||||||||
Habitat | Rocky or gravelly slopes | |||||||||||||||||||||||||||||
Elevation | 2100-3400 m (6900-11200 ft) | |||||||||||||||||||||||||||||
Distribution |
AK; AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NL; NS; NT; ON; PE; QC; SK; YT; nearly worldwide
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CA; NV; OR
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Discussion | Subspecies 7+ (6 in the flora). Polygonum aviculare is a taxonomically controversial polyploid complex of selfing annuals. Although members of the complex have been considered inbreeders, they possess some structures that make cross pollination possible. Cleistogamous and chasmogamous flowers, heterostyly, protandry, and the capacity to secrete nectar suggest an ancestral mixed-mating system. Isoenzyme studies showed that the complex has an allopolyploid origin (P. Meerts et al. 1998) and has evolved as a swarm of inbreeding lines (“Jordanons”) (J. Gasquez et al. 1978). The six subspecies included here have been treated variously (T. Karlsson 2000; M. Costea and F. J. Tardif 2003). Complex intergradation patterns among them make their recognition at the species level impractical. Multivariate analysis and isoenzyme studies show that populations with intermediate characteristics may occur (Meerts et al. 1990, 1998). Except for subsp. boreale, which occurs in Greenland and Labrador, all subspecies are partially sympatric and their distributions have been influenced greatly by humans. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Polygonum shastense may be cultivated in rock gardens in rock crevices with favorable water regime and shaded in summer. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 5, p. 556. | FNA vol. 5, p. 563. | ||||||||||||||||||||||||||||
Parent taxa | Polygonaceae > subfam. Polygonoideae > Polygonum > sect. Polygonum | Polygonaceae > subfam. Polygonoideae > Polygonum > sect. Duravia | ||||||||||||||||||||||||||||
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Name authority | Linnaeus: Sp. Pl. 1: 362. (1753) | W. H. Brewer: Proc. Amer. Acad. Arts 8: 400. (1872) | ||||||||||||||||||||||||||||
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