Polygonum aviculare |
Polygonum ramosissimum |
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birdweed, common knotgrass, common knotweed, doorweed, dooryard knotweed, knotgrass, knotweed, lowgrass, pigweed, prostrate knotweed, renouée des oiseaux, yard knotweed |
bushy knotweed, yellow knotweed, yellow-flower knotweed |
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Habit | Plants green or bluish green, green after drying, sometimes whitish from powdery mildew, homophyllous or heterophyllous. | Plants yellowish green or bluish green, (dark brown to black after drying in subsp. prolificum), heterophyllous or homophyllous. | ||||||||||||||||||||||||||||||||
Stems | prostrate to erect, branched, flexuous, 5–200 cm. |
erect, usually profusely branched in distal 1/2, not wiry, 10–100(–200) cm. |
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Leaves | ocrea 3–15 mm, proximal part cylindric or ± funnelform, distal part silvery, hyaline, soon disintegrating into persistent fibers or nearly completely deciduous; petiole 0.3–9 mm; blade green to gray-green, narrowly elliptic, lanceolate, elliptic, obovate, or spatulate, 6–50(–60) × 0.5–22 mm, margins flat, apex acute, obtuse, or rounded; stem leaves 1–4 times as long as adjacent branch leaves; distal leaves overtopping flowers. |
ocrea 6–12(–15) mm, proximal part cylindric, distal part silvery, soon disintegrating into persistent brown fibers; petiole 2–4 mm; blade variable, light yellowish green to bluish green, proximal often caducous, narrowly elliptic, lanceolate, or oblanceolate, rarely ovate, 8–70 × 4–18(–35) mm, margins flat, apex acute to acuminate or obtuse; distal leaves, either overtopping or shorter than or equaling flowers. |
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Inflorescences | axillary; cymes uniformly distributed or aggregated at tips of stems and branches, 1–6(–8)-flowered. |
axillary or axillary and terminal, spikelike; cymes uniformly distributed or crowded toward tips of branches, 2–5-flowered. |
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Pedicels | enclosed in or exserted from ocreae, 1.5–5 mm. |
enclosed in or exserted from ocreae, 1–6 mm. |
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Flowers | closed or semi-open; perianth 1.8–5.5 mm; tube 20–57% of perianth length; tepals overlapping or not, green or reddish brown with white, pink, or red margins, petaloid, not keeled, oblong to obovate, often cucullate in fruit; midveins branched or unbranched, thickened or not; stamens 5–8. |
closed; perianth (2–)2.2–3.6(–4) mm; tube 20–38% of perianth length; tepals overlapping, greenish yellow with greenish yellow or yellow, rarely pink or white, margins, petaloid or sepaloid, not keeled, elliptic to oblong, cucullate; midveins thickened or not; stamens 3–6(–8). |
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Achenes | enclosed in or exserted from perianth, light to dark brown, ovate, (2–)3-gonous, 1.2–4.2 mm, faces subequal or unequal, apex not beaked, edges slightly concave, dull, usually coarsely striate-tubercled, sometimes obscurely tubercled; late-season achenes common or not, 2–5 mm. |
enclosed in or exserted from perianth, dark brown, ovate, 3-gonous, 1.6–3.5 mm, faces subequal, concave, apex not beaked, edges straight, shiny or dull, usually smooth to roughened, sometimes uniformly or obscurely tubercled; late-season achenes common, 4–15 mm. |
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Polygonum aviculare |
Polygonum ramosissimum |
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Distribution |
AK; AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NL; NS; NT; ON; PE; QC; SK; YT; nearly worldwide
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AR; AZ; CA; CO; CT; DE; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NS; NT; ON; PE; QC; SK
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Discussion | Subspecies 7+ (6 in the flora). Polygonum aviculare is a taxonomically controversial polyploid complex of selfing annuals. Although members of the complex have been considered inbreeders, they possess some structures that make cross pollination possible. Cleistogamous and chasmogamous flowers, heterostyly, protandry, and the capacity to secrete nectar suggest an ancestral mixed-mating system. Isoenzyme studies showed that the complex has an allopolyploid origin (P. Meerts et al. 1998) and has evolved as a swarm of inbreeding lines (“Jordanons”) (J. Gasquez et al. 1978). The six subspecies included here have been treated variously (T. Karlsson 2000; M. Costea and F. J. Tardif 2003). Complex intergradation patterns among them make their recognition at the species level impractical. Multivariate analysis and isoenzyme studies show that populations with intermediate characteristics may occur (Meerts et al. 1990, 1998). Except for subsp. boreale, which occurs in Greenland and Labrador, all subspecies are partially sympatric and their distributions have been influenced greatly by humans. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Subspecies 2 (2 in the flora). Polygonum ramosissimum exhibits considerable morphological complexity and is similar in difficulty to the P. aviculare complex. Further research is necessary to understand the infraspecific variability of this species (M. Costea and F. J. Tardif 2003b). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 5, p. 556. | FNA vol. 5, p. 551. | ||||||||||||||||||||||||||||||||
Parent taxa | Polygonaceae > subfam. Polygonoideae > Polygonum > sect. Polygonum | Polygonaceae > subfam. Polygonoideae > Polygonum > sect. Polygonum | ||||||||||||||||||||||||||||||||
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Name authority | Linnaeus: Sp. Pl. 1: 362. (1753) | Michaux: Fl. Bor.-Amer. 1: 237. (1803) | ||||||||||||||||||||||||||||||||
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