Polygonum aviculare |
Polygonum paronychia |
|||||||||||||||||||||||||||||
---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
birdweed, common knotgrass, common knotweed, doorweed, dooryard knotweed, knotgrass, knotweed, lowgrass, pigweed, prostrate knotweed, renouée des oiseaux, yard knotweed |
beach knotweed, beach or black or dune knotweed, black knotweed, dune knotweed |
|||||||||||||||||||||||||||||
Habit | Plants green or bluish green, green after drying, sometimes whitish from powdery mildew, homophyllous or heterophyllous. | Shrubs or subshrubs. | ||||||||||||||||||||||||||||
Stems | prostrate to erect, branched, flexuous, 5–200 cm. |
prostrate or ascending, brown, branched, rooting at nodes, not wiry, 10–100 cm, glabrous, covered with remains of lacerate, hyaline ocreae. |
||||||||||||||||||||||||||||
Leaves | ocrea 3–15 mm, proximal part cylindric or ± funnelform, distal part silvery, hyaline, soon disintegrating into persistent fibers or nearly completely deciduous; petiole 0.3–9 mm; blade green to gray-green, narrowly elliptic, lanceolate, elliptic, obovate, or spatulate, 6–50(–60) × 0.5–22 mm, margins flat, apex acute, obtuse, or rounded; stem leaves 1–4 times as long as adjacent branch leaves; distal leaves overtopping flowers. |
crowded at branch tips, articulated to ocreae, basal leaves caducous or persistent, distal leaves not reduced in size; ocrea 15–20 mm, glabrous, proximal part cylindric to funnelform, distal part silvery, entire or slightly lacerate, disintegrating into persistent white-gray curly fibers; petiole 0–0.5 mm; blade 1-veined, without pleats, linear to oblanceolate, (5–)10–20(–33) × 3–8 mm, coriaceous, margins revolute, smooth, apex acute or mucronate. |
||||||||||||||||||||||||||||
Inflorescences | axillary; cymes uniformly distributed or aggregated at tips of stems and branches, 1–6(–8)-flowered. |
axillary; cymes crowded in distal axils, 2–5-flowered. |
||||||||||||||||||||||||||||
Pedicels | enclosed in or exserted from ocreae, 1.5–5 mm. |
enclosed in ocreae, erect to spreading, 2–5 mm. |
||||||||||||||||||||||||||||
Flowers | closed or semi-open; perianth 1.8–5.5 mm; tube 20–57% of perianth length; tepals overlapping or not, green or reddish brown with white, pink, or red margins, petaloid, not keeled, oblong to obovate, often cucullate in fruit; midveins branched or unbranched, thickened or not; stamens 5–8. |
semi-open or open; perianth (4.5–)6–10 mm; tube 22–48% of perianth length; tepals partially overlapping, uniformly pink or white, reddish brown when dried, petaloid, oblong-ovate to ± lanceolate, apex rounded; midveins pinnately branched; stamens 8. |
||||||||||||||||||||||||||||
Achenes | enclosed in or exserted from perianth, light to dark brown, ovate, (2–)3-gonous, 1.2–4.2 mm, faces subequal or unequal, apex not beaked, edges slightly concave, dull, usually coarsely striate-tubercled, sometimes obscurely tubercled; late-season achenes common or not, 2–5 mm. |
enclosed in or slightly exserted from perianth, black, ovate, 4–5 mm, faces subequal, shiny, smooth. |
||||||||||||||||||||||||||||
Polygonum aviculare |
Polygonum paronychia |
|||||||||||||||||||||||||||||
Phenology | Flowering Mar–Sep. | |||||||||||||||||||||||||||||
Habitat | Coastal sands, scrub along coast | |||||||||||||||||||||||||||||
Elevation | 0-50 m (0-200 ft) | |||||||||||||||||||||||||||||
Distribution |
AK; AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NL; NS; NT; ON; PE; QC; SK; YT; nearly worldwide
|
CA; OR; WA; BC
|
||||||||||||||||||||||||||||
Discussion | Subspecies 7+ (6 in the flora). Polygonum aviculare is a taxonomically controversial polyploid complex of selfing annuals. Although members of the complex have been considered inbreeders, they possess some structures that make cross pollination possible. Cleistogamous and chasmogamous flowers, heterostyly, protandry, and the capacity to secrete nectar suggest an ancestral mixed-mating system. Isoenzyme studies showed that the complex has an allopolyploid origin (P. Meerts et al. 1998) and has evolved as a swarm of inbreeding lines (“Jordanons”) (J. Gasquez et al. 1978). The six subspecies included here have been treated variously (T. Karlsson 2000; M. Costea and F. J. Tardif 2003). Complex intergradation patterns among them make their recognition at the species level impractical. Multivariate analysis and isoenzyme studies show that populations with intermediate characteristics may occur (Meerts et al. 1990, 1998). Except for subsp. boreale, which occurs in Greenland and Labrador, all subspecies are partially sympatric and their distributions have been influenced greatly by humans. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Polygonum paronychia may be cultivated in rock gardens in open sites with sandy soil. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
||||||||||||||||||||||||||||
Key |
|
|||||||||||||||||||||||||||||
Source | FNA vol. 5, p. 556. | FNA vol. 5, p. 562. | ||||||||||||||||||||||||||||
Parent taxa | Polygonaceae > subfam. Polygonoideae > Polygonum > sect. Polygonum | Polygonaceae > subfam. Polygonoideae > Polygonum > sect. Duravia | ||||||||||||||||||||||||||||
Sibling taxa | ||||||||||||||||||||||||||||||
Subordinate taxa | ||||||||||||||||||||||||||||||
Name authority | Linnaeus: Sp. Pl. 1: 362. (1753) | Chamisso & Schlechtendal: Linnaea 3: 51. (1828) | ||||||||||||||||||||||||||||
Web links |
|
|