Polygonum aviculare |
Polygonum heterosepalum |
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birdweed, common knotgrass, common knotweed, doorweed, dooryard knotweed, knotgrass, knotweed, lowgrass, pigweed, prostrate knotweed, renouée des oiseaux, yard knotweed |
dwarf desert knotweed, odd-sepal knotweed |
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Habit | Plants green or bluish green, green after drying, sometimes whitish from powdery mildew, homophyllous or heterophyllous. | Herbs, compact, often cushion-like. | ||||||||||||||||||||||||||||
Stems | prostrate to erect, branched, flexuous, 5–200 cm. |
erect, green or reddish, simple or branched near base, not wiry, 1.5–5 cm, glabrous. |
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Leaves | ocrea 3–15 mm, proximal part cylindric or ± funnelform, distal part silvery, hyaline, soon disintegrating into persistent fibers or nearly completely deciduous; petiole 0.3–9 mm; blade green to gray-green, narrowly elliptic, lanceolate, elliptic, obovate, or spatulate, 6–50(–60) × 0.5–22 mm, margins flat, apex acute, obtuse, or rounded; stem leaves 1–4 times as long as adjacent branch leaves; distal leaves overtopping flowers. |
uniformly distributed, dense, not articulated to ocreae, basal leaves persistent or caducous, distal leaves gradually reduced to bracts; ocrea 3–6 mm, glabrous, proximal part cylindric, distal part disintegrating almost to base, with whitish, straight, rigid fibers; petiole absent; blade 3-veined, without pleats, linear to lanceolate, 10–20 × 0.6–2.7 mm, margins revolute, smooth, apex spine-tipped. |
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Inflorescences | axillary; cymes uniformly distributed or aggregated at tips of stems and branches, 1–6(–8)-flowered. |
axillary; cymes in most axils, 2–3-flowered. |
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Pedicels | enclosed in or exserted from ocreae, 1.5–5 mm. |
enclosed in ocreae, erect, 0.1–1 mm. |
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Flowers | closed or semi-open; perianth 1.8–5.5 mm; tube 20–57% of perianth length; tepals overlapping or not, green or reddish brown with white, pink, or red margins, petaloid, not keeled, oblong to obovate, often cucullate in fruit; midveins branched or unbranched, thickened or not; stamens 5–8. |
closed; perianth 2.3–2.7 mm; tube 3–7% of perianth length; tepals overlapping, whitish with whitish or pink margins, petaloid, oblong, navicular, dimorphic, outer 2 shorter than inner 3, outer 2 0.8–1.2 mm, inner 2.3–2.7 mm, papillose at base, apex acute or acuminate; midveins unbranched; stamens 5–6. |
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Achenes | enclosed in or exserted from perianth, light to dark brown, ovate, (2–)3-gonous, 1.2–4.2 mm, faces subequal or unequal, apex not beaked, edges slightly concave, dull, usually coarsely striate-tubercled, sometimes obscurely tubercled; late-season achenes common or not, 2–5 mm. |
enclosed in perianth, olive brown, narrowly ovate to ovate-lanceolate, 1.5–2 mm, faces subequal, shiny, smooth. |
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Polygonum aviculare |
Polygonum heterosepalum |
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Phenology | Flowering Jun–Aug. | |||||||||||||||||||||||||||||
Habitat | Dry waste ground, open flats in sagebrush plains, ponderosa pine forests | |||||||||||||||||||||||||||||
Elevation | 1000-1500 m (3300-4900 ft) | |||||||||||||||||||||||||||||
Distribution |
AK; AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NL; NS; NT; ON; PE; QC; SK; YT; nearly worldwide
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ID; NV; OR |
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Discussion | Subspecies 7+ (6 in the flora). Polygonum aviculare is a taxonomically controversial polyploid complex of selfing annuals. Although members of the complex have been considered inbreeders, they possess some structures that make cross pollination possible. Cleistogamous and chasmogamous flowers, heterostyly, protandry, and the capacity to secrete nectar suggest an ancestral mixed-mating system. Isoenzyme studies showed that the complex has an allopolyploid origin (P. Meerts et al. 1998) and has evolved as a swarm of inbreeding lines (“Jordanons”) (J. Gasquez et al. 1978). The six subspecies included here have been treated variously (T. Karlsson 2000; M. Costea and F. J. Tardif 2003). Complex intergradation patterns among them make their recognition at the species level impractical. Multivariate analysis and isoenzyme studies show that populations with intermediate characteristics may occur (Meerts et al. 1990, 1998). Except for subsp. boreale, which occurs in Greenland and Labrador, all subspecies are partially sympatric and their distributions have been influenced greatly by humans. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 5, p. 556. | FNA vol. 5, p. 563. | ||||||||||||||||||||||||||||
Parent taxa | Polygonaceae > subfam. Polygonoideae > Polygonum > sect. Polygonum | Polygonaceae > subfam. Polygonoideae > Polygonum > sect. Duravia | ||||||||||||||||||||||||||||
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Name authority | Linnaeus: Sp. Pl. 1: 362. (1753) | M. Peck & Ownbey: Madroño 10: 250. (1950) | ||||||||||||||||||||||||||||
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