Polygonum aviculare |
Polygonum fowleri |
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birdweed, common knotgrass, common knotweed, doorweed, dooryard knotweed, knotgrass, knotweed, lowgrass, pigweed, prostrate knotweed, renouée des oiseaux, yard knotweed |
Fowler's knotweed, Hudsonian knotweed |
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Habit | Plants green or bluish green, green after drying, sometimes whitish from powdery mildew, homophyllous or heterophyllous. | Plants green, sometimes purple tinged, homophyllous or heterophyllous, sometimes subsucculent. | ||||||||||||||||||||||||||||||||
Stems | prostrate to erect, branched, flexuous, 5–200 cm. |
prostrate to ascending, sometimes zigzagged, branched from base, not wiry, 5–50 cm. |
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Leaves | ocrea 3–15 mm, proximal part cylindric or ± funnelform, distal part silvery, hyaline, soon disintegrating into persistent fibers or nearly completely deciduous; petiole 0.3–9 mm; blade green to gray-green, narrowly elliptic, lanceolate, elliptic, obovate, or spatulate, 6–50(–60) × 0.5–22 mm, margins flat, apex acute, obtuse, or rounded; stem leaves 1–4 times as long as adjacent branch leaves; distal leaves overtopping flowers. |
ocrea 2.5–12 mm, proximal part funnelform, distal part soon disintegrating, nearly completely deciduous or fibers persistent; petiole 2–7 mm; blade light green, sometimes purple tinged, elliptic to elliptic-obovate or obovate, 8–30(–50) × 4–15(–25) mm, margins flat, apex acute to obtuse; middle stem leaves 1.1–2.1(–3.4) times as long as adjacent branch leaves, distal leaves overtopping flowers. |
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Inflorescences | axillary; cymes uniformly distributed or aggregated at tips of stems and branches, 1–6(–8)-flowered. |
axillary; cymes uniformly distributed, 1–7(–10)-flowered. |
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Pedicels | enclosed in or exserted from ocreae, 1.5–5 mm. |
enclosed in or sometimes exserted from ocreae, 1–2.5 mm. |
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Flowers | closed or semi-open; perianth 1.8–5.5 mm; tube 20–57% of perianth length; tepals overlapping or not, green or reddish brown with white, pink, or red margins, petaloid, not keeled, oblong to obovate, often cucullate in fruit; midveins branched or unbranched, thickened or not; stamens 5–8. |
closed; perianth (2.2–)2.5–4.5 mm; tube 23–38% of perianth length; tepals initially overlapping, pushed apart as achene develops, green, margins white to pink, petaloid, not keeled, oblong, cucullate; midveins branched, sometimes not visible; stamens 6–8. |
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Achenes | enclosed in or exserted from perianth, light to dark brown, ovate, (2–)3-gonous, 1.2–4.2 mm, faces subequal or unequal, apex not beaked, edges slightly concave, dull, usually coarsely striate-tubercled, sometimes obscurely tubercled; late-season achenes common or not, 2–5 mm. |
exserted from perianth, brown to dark brown, broadly ovate to ovate-lanceolate, (2–)3-gonous, (1.8–)2–3.7(–4.5) mm, faces subequal or unequal, flat to concave, apex beaked, edges strongly concave, shiny to dull, roughened, rarely obscurely tubercled; late-season achenes common, 4–6 mm. |
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Polygonum aviculare |
Polygonum fowleri |
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Distribution |
AK; AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NL; NS; NT; ON; PE; QC; SK; YT; nearly worldwide
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AK; CA; ME; OR; WA; BC; MB; NB; NS; NT; NU; ON; PE; QC; YT
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Discussion | Subspecies 7+ (6 in the flora). Polygonum aviculare is a taxonomically controversial polyploid complex of selfing annuals. Although members of the complex have been considered inbreeders, they possess some structures that make cross pollination possible. Cleistogamous and chasmogamous flowers, heterostyly, protandry, and the capacity to secrete nectar suggest an ancestral mixed-mating system. Isoenzyme studies showed that the complex has an allopolyploid origin (P. Meerts et al. 1998) and has evolved as a swarm of inbreeding lines (“Jordanons”) (J. Gasquez et al. 1978). The six subspecies included here have been treated variously (T. Karlsson 2000; M. Costea and F. J. Tardif 2003). Complex intergradation patterns among them make their recognition at the species level impractical. Multivariate analysis and isoenzyme studies show that populations with intermediate characteristics may occur (Meerts et al. 1990, 1998). Except for subsp. boreale, which occurs in Greenland and Labrador, all subspecies are partially sympatric and their distributions have been influenced greatly by humans. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Subspecies 2 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 5, p. 556. | FNA vol. 5, p. 554. | ||||||||||||||||||||||||||||||||
Parent taxa | Polygonaceae > subfam. Polygonoideae > Polygonum > sect. Polygonum | Polygonaceae > subfam. Polygonoideae > Polygonum > sect. Polygonum | ||||||||||||||||||||||||||||||||
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Name authority | Linnaeus: Sp. Pl. 1: 362. (1753) | B. L. Robinson: Rhodora 4: 67, plate 35, figs. 14, 15. (1902) | ||||||||||||||||||||||||||||||||
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