Polygonum aviculare |
Polygonum bolanderi |
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birdweed, common knotgrass, common knotweed, doorweed, dooryard knotweed, knotgrass, knotweed, lowgrass, pigweed, prostrate knotweed, renouée des oiseaux, yard knotweed |
Bolander's knotweed, Bolander's polygonum |
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Habit | Plants green or bluish green, green after drying, sometimes whitish from powdery mildew, homophyllous or heterophyllous. | Shrubs. | ||||||||||||||||||||||||||||
Stems | prostrate to erect, branched, flexuous, 5–200 cm. |
erect, brown, simple, wiry, gnarled with age, 20–60 cm, glabrous. |
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Leaves | ocrea 3–15 mm, proximal part cylindric or ± funnelform, distal part silvery, hyaline, soon disintegrating into persistent fibers or nearly completely deciduous; petiole 0.3–9 mm; blade green to gray-green, narrowly elliptic, lanceolate, elliptic, obovate, or spatulate, 6–50(–60) × 0.5–22 mm, margins flat, apex acute, obtuse, or rounded; stem leaves 1–4 times as long as adjacent branch leaves; distal leaves overtopping flowers. |
crowded at branch tips, not articulated to ocreae, basal leaves caducous or persistent, distal leaves abruptly reduced to bracts; ocrea 6–10 mm, glabrous or papillose-scabridulous, proximal part cylindric, distal part silvery, deeply fringed, disintegrating; petiole absent; blade 3-veined, without pleats, linear to subulate, 3–15(–25) × 0.4–1.5 mm, margins flat, smooth, apex acuminate to spine-tipped. |
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Inflorescences | axillary; cymes uniformly distributed or aggregated at tips of stems and branches, 1–6(–8)-flowered. |
axillary; cymes in distal axils, 1–2-flowered. |
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Pedicels | enclosed in or exserted from ocreae, 1.5–5 mm. |
absent. |
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Flowers | closed or semi-open; perianth 1.8–5.5 mm; tube 20–57% of perianth length; tepals overlapping or not, green or reddish brown with white, pink, or red margins, petaloid, not keeled, oblong to obovate, often cucullate in fruit; midveins branched or unbranched, thickened or not; stamens 5–8. |
semi-open; perianth 2.6–3.2 mm; tube 18–33% of perianth length; tepals overlapping, ± recurved, uniformly white to pink, petaloid, elliptic-oblong to obovate, navicular, apex rounded; midveins unbranched; stamens 8. |
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Achenes | enclosed in or exserted from perianth, light to dark brown, ovate, (2–)3-gonous, 1.2–4.2 mm, faces subequal or unequal, apex not beaked, edges slightly concave, dull, usually coarsely striate-tubercled, sometimes obscurely tubercled; late-season achenes common or not, 2–5 mm. |
enclosed in perianth, light brown, lanceolate to oblong-ovate, 2.5–3 mm, faces subequal, shiny, smooth. |
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Polygonum aviculare |
Polygonum bolanderi |
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Phenology | Flowering Jun–Nov. | |||||||||||||||||||||||||||||
Habitat | Open, dry, gravelly, rocky places | |||||||||||||||||||||||||||||
Elevation | 300-1500 m (1000-4900 ft) | |||||||||||||||||||||||||||||
Distribution |
AK; AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NL; NS; NT; ON; PE; QC; SK; YT; nearly worldwide
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CA |
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Discussion | Subspecies 7+ (6 in the flora). Polygonum aviculare is a taxonomically controversial polyploid complex of selfing annuals. Although members of the complex have been considered inbreeders, they possess some structures that make cross pollination possible. Cleistogamous and chasmogamous flowers, heterostyly, protandry, and the capacity to secrete nectar suggest an ancestral mixed-mating system. Isoenzyme studies showed that the complex has an allopolyploid origin (P. Meerts et al. 1998) and has evolved as a swarm of inbreeding lines (“Jordanons”) (J. Gasquez et al. 1978). The six subspecies included here have been treated variously (T. Karlsson 2000; M. Costea and F. J. Tardif 2003). Complex intergradation patterns among them make their recognition at the species level impractical. Multivariate analysis and isoenzyme studies show that populations with intermediate characteristics may occur (Meerts et al. 1990, 1998). Except for subsp. boreale, which occurs in Greenland and Labrador, all subspecies are partially sympatric and their distributions have been influenced greatly by humans. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Polygonum bolanderi is known from several counties in northwestern California (Butte, Napa, Shasta, and Sonoma). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 5, p. 556. | FNA vol. 5, p. 564. | ||||||||||||||||||||||||||||
Parent taxa | Polygonaceae > subfam. Polygonoideae > Polygonum > sect. Polygonum | Polygonaceae > subfam. Polygonoideae > Polygonum > sect. Duravia | ||||||||||||||||||||||||||||
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Synonyms | Duravia bolanderi | |||||||||||||||||||||||||||||
Name authority | Linnaeus: Sp. Pl. 1: 362. (1753) | W. H. Brewer: Proc. Amer. Acad. Arts 8: 400. (1872) | ||||||||||||||||||||||||||||
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