Polygonum achoreum |
Polygonum aviculare |
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beak-seed knotweed, Blake's knotweed, leathery knotweed, renouée coriace, striate knotweed |
birdweed, common knotgrass, common knotweed, doorweed, dooryard knotweed, knotgrass, knotweed, lowgrass, pigweed, prostrate knotweed, renouée des oiseaux, yard knotweed |
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Habit | Plants light green (often covered with whitish powdery mildew), homophyllous or, sometimes, heterophyllous. | Plants green or bluish green, green after drying, sometimes whitish from powdery mildew, homophyllous or heterophyllous. | ||||||||||||||||||||||||||||
Stems | erect when young, decumbent or prostrate later, moderately branched especially from base, not wiry, 50–70 cm. |
prostrate to erect, branched, flexuous, 5–200 cm. |
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Leaves | ocreae 5–12 mm, proximal part cylindric, distal part soon disintegrating into brown fibers; petiole 0.3–1.5 mm; blade light yellowish green, elliptic to obovate, 8–35 × 3–15 mm, margins flat, apex rounded; stem leaves 1–2.1(–3) times longer than branch leaves; distal leaves overtopping flowers. |
ocrea 3–15 mm, proximal part cylindric or ± funnelform, distal part silvery, hyaline, soon disintegrating into persistent fibers or nearly completely deciduous; petiole 0.3–9 mm; blade green to gray-green, narrowly elliptic, lanceolate, elliptic, obovate, or spatulate, 6–50(–60) × 0.5–22 mm, margins flat, apex acute, obtuse, or rounded; stem leaves 1–4 times as long as adjacent branch leaves; distal leaves overtopping flowers. |
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Inflorescences | axillary, cymes in axils of most leaves and toward tips of stems and branchs, 1–3(–5)-flowered. |
axillary; cymes uniformly distributed or aggregated at tips of stems and branches, 1–6(–8)-flowered. |
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Pedicels | enclosed in ocreae, 1.3–1.8(–2) mm. |
enclosed in or exserted from ocreae, 1.5–5 mm. |
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Flowers | closed; perianth 2.6–4 mm; tube 40–55% of perianth length; tepals incurved, yellow-green with yellow to green, rarely pinkish, margins, sepaloid, ± keeled, narrowly oblong, cucullate; midveins unbranched, moderately to heavily thickened, tepals appearing keeled; stamens 5–8. |
closed or semi-open; perianth 1.8–5.5 mm; tube 20–57% of perianth length; tepals overlapping or not, green or reddish brown with white, pink, or red margins, petaloid, not keeled, oblong to obovate, often cucullate in fruit; midveins branched or unbranched, thickened or not; stamens 5–8. |
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Achenes | enclosed in perianth, yellow-green to tan, ovate, 3-gonous, 2.4–3.5 mm, faces unequal, apex not beaked, edges concave or nearly straight, dull, uniformly tubercled; late-season achenes common, 3–5 mm. |
enclosed in or exserted from perianth, light to dark brown, ovate, (2–)3-gonous, 1.2–4.2 mm, faces subequal or unequal, apex not beaked, edges slightly concave, dull, usually coarsely striate-tubercled, sometimes obscurely tubercled; late-season achenes common or not, 2–5 mm. |
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2n | = 40, 60. |
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Polygonum achoreum |
Polygonum aviculare |
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Phenology | Flowering Jul–Sep. | |||||||||||||||||||||||||||||
Habitat | Disturbed areas, roadsides, sidewalks, edges of cultivated fields | |||||||||||||||||||||||||||||
Elevation | 10-800 m (0-2600 ft) | |||||||||||||||||||||||||||||
Distribution |
AK; CO; CT; IA; ID; IL; IN; KS; ME; MI; MN; MO; MT; ND; NE; NV; NY; OH; OR; SD; UT; VT; WA; WI; WV; WY; AB; BC; MB; NB; NS; NT; ON; QC; SK; YT
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AK; AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NL; NS; NT; ON; PE; QC; SK; YT; nearly worldwide
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Discussion | Polygonum achoreum frequently is confused with P. erectum. It can be distinguished by its usually homophyllous leaves, its perianth, which is enlarged at the base and constricted above the fruit, its longer perianth tube, and its yellow-green to tan, tubercled achenes. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Subspecies 7+ (6 in the flora). Polygonum aviculare is a taxonomically controversial polyploid complex of selfing annuals. Although members of the complex have been considered inbreeders, they possess some structures that make cross pollination possible. Cleistogamous and chasmogamous flowers, heterostyly, protandry, and the capacity to secrete nectar suggest an ancestral mixed-mating system. Isoenzyme studies showed that the complex has an allopolyploid origin (P. Meerts et al. 1998) and has evolved as a swarm of inbreeding lines (“Jordanons”) (J. Gasquez et al. 1978). The six subspecies included here have been treated variously (T. Karlsson 2000; M. Costea and F. J. Tardif 2003). Complex intergradation patterns among them make their recognition at the species level impractical. Multivariate analysis and isoenzyme studies show that populations with intermediate characteristics may occur (Meerts et al. 1990, 1998). Except for subsp. boreale, which occurs in Greenland and Labrador, all subspecies are partially sympatric and their distributions have been influenced greatly by humans. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 5, p. 551. | FNA vol. 5, p. 556. | ||||||||||||||||||||||||||||
Parent taxa | Polygonaceae > subfam. Polygonoideae > Polygonum > sect. Polygonum | Polygonaceae > subfam. Polygonoideae > Polygonum > sect. Polygonum | ||||||||||||||||||||||||||||
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Synonyms | P. erectum subsp. achoreum | |||||||||||||||||||||||||||||
Name authority | S. F. Blake: Rhodora 19: 232. (1917) | Linnaeus: Sp. Pl. 1: 362. (1753) | ||||||||||||||||||||||||||||
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