Polyctenium
Polyctenium fremontii
comb-leaf
combleaf, desert comb-leaf, Fremont's polyctenium, two-grooved comb-leaf, Washoe comb-leaf
erect to ascending, branched distally.
few to numerous from base, (0.2–)0.5–1.6(–2) dm, usually sparsely to densely pubescent proximally, sometimes glaucous and glabrous distally.
cauline;
petiolate or subsessile;
blade (base not auriculate), not rosulate;
margins pinnatifid.
often persistent in subsequent seasons as pectinate spines;
petiolate proximally, subsessile distally;
petiole winged, 0–3(–5) mm;
blade linear (in outline), (0.5–)1–2.2(–2.8) cm, rigid, surfaces sparsely to densely pubescent, trichomes dendritic, margin and apex often with simple or forked ones;
lobes (2–)3–4(–5) each side, with prominent midvein and narrow band of blade;
terminal lobe (0.2–)0.4–0.7(–1) cm × (0.3–)0.5–1(–1.5) mm, as long as or slightly longer than adjacent lateral lobes.
(corymbose, several-flowered), congested or considerably elongated in fruit.
sepals (erect or slightly ascending), oblong-ovate;
petals white or purplish, obovate to oblanceolate, (longer than sepals, claw undifferentiated from blade, apex obtuse);
stamens slightly tetradynamous;
filaments not dilated basally;
anthers ovate, (apex obtuse);
nectar glands confluent, subtending bases of stamens.
sepals 1.5–2 × 0.9–1.1 mm, hyaline-margined, glabrous or sparsely pubescent;
petals 3–4 × 1.5–2 mm, attenuate to base;
filaments 1.2–2 mm;
anthers 0.4–0.5 mm.
divaricate-ascending, slender.
(2–)4–7(–10) mm, sparsely pubescent or glabrous.
(siliques or silicles), sessile, usually linear to oblong, rarely ovate-oblong, not torulose, slightly to strongly angustiseptate;
valves each not veined, glabrous or, rarely, sparsely puberulent;
replum rounded;
septum complete;
ovules 30–46 per ovary;
stigma capitate, entire.
erect to ascending, (0.2–)0.4–1.3(–1.8) cm × 0.9–2 mm;
style (0.2–)0.4–0.7(–1) mm.
uniseriate, plump, not winged, oblong;
seed coat not mucilaginous when wetted;
cotyledons incumbent.
0.7–0.9 × 0.3–0.5 mm.
Polyctenium
Polyctenium fremontii
Species 1.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Polyctenium fremontii is highly variable in fruit size and the compactness of the fruiting raceme, but in habit, flower size and color, leaf morphology, indumentum, fruiting pedicel length and orientation, number of ovules per ovary, and basically every other aspect of the plants, it is quite constant. If one examines only the types of those two taxa and that of P. fremontii, it seems that perhaps two or three taxa might be recognized. Upon careful study of extensive material, one realizes that only one taxon, instead of three or more, is represented. The alleged differences between P. fremontii and P. williamsiae in characters other than fruit morphology do not hold. As for fruit size, it was said to be 2–4 × 2–2.5 mm in P. williamsiae and (4–)6–13(–20) × 1–2 mm in P. fremontii. Fruit lengths in material annotated by Rollins as P. fremontii are 2–7 mm in Tiehm 8108 and 3.5–11 mm in Ertter 5726, both at GH. Furthermore, the compactness of the infructescence can be equally variable, and in the holotype of var. confertum there are 12–15 pedicels along 1 cm in the middle of the rachis, whereas in Ertter 5726 (GH) there are 6–12. On one sheet, Schoolcraft 1287 (GH), compact and lax racemes and relatively short (3 mm) and longer (7 mm) fruits are represented. The variation in fruit length and width depends largely upon the number of ovules maturing into seeds, and in plants with very short fruits, including the type collection of P. williamsiae, none of the ovules matured into seeds, whereas in those with longest and narrowest fruits almost all ovules matured into seeds. Regardless of how long the fruit is or how many ovules mature into seeds, the ovule number is fairly constant throughout the range of the species. In my opinion, except for the type species of Polyctenium, all of the other taxa recognized in this genus do not represent biologically distinct entities.
To my knowledge, Polyctenium fremontii is known from counties in California (Lassen, Modoc, Mono, Siskiyou), Idaho (Gooding), Nevada (Churchill, Douglas, Humboldt, Lyon, Mineral, Washoe), and Oregon (Crook, Deschutes, Harney, Klamath, Lake, Malheur).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
- Local floras:
CA, 
OR, 
WA - Local Web sites: CalFlora, CalPhotos,
Flora NW, 
PNW Herbaria 
WildflowerSearch
iNaturalist (observations)
USDA Plants Database- LBJ Wildflower Center
- SEINet
- Plants of the World Online
- Encyclopedia of Life
- Wikipedia
- Google Image Search
- Local floras: CA, OR, WA
- Local Web sites: CalFlora, CalPhotos, Flora NW, PNW Herbaria
- WildflowerSearch
- iNaturalist (observations)
- USDA Plants Database
- LBJ Wildflower Center
- SEINet
- Plants of the World Online
- Encyclopedia of Life
- Wikipedia
- Google Image Search
