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pale nodding-cap moss, pale pohlia, Wahlenberg's pohlia moss

ludwig's pohlia moss, ludwig's thread-moss

Habit Plants small to very large, whitish or sometimes in large forms reddish, dull. Plants medium-sized to large, green, sometimes red tinged, dull.
Stems

0.8–10 cm.

0.8–3.5 cm.

Leaves

laxly spreading to more stiffly erect, lanceolate to ovate-lanceolate, 0.6–1.2 mm;

margins weakly to strongly serrulate in distal 1/3;

costa ending well before apex;

distal medial laminal cells laxly and broadly hexagonal to rhomboidal, 65–110 µm, walls thin.

erect to ± spreading, lanceolate to broadly lanceolate, 1.2–2.4 mm;

base long-decurrent;

margins serrulate to serrate in distal 1/3;

costa subpercurrent;

distal medial laminal cells rhombic to rhomboidal, 40–80 µm, walls firm to thin.

Seta

orange-brown.

orange-brown.

Sexual condition

dioicous;

perigonial leaves ovate;

perichaetial leaves weakly differentiated, lanceolate.

dioicous;

perigonial leaves broadly short-ovate;

perichaetial leaves scarcely differentiated, broadly lanceolate.

Capsule

inclined ± 180°, brown to red-brown, sometimes stramineous, short-pyriform to urceolate, neck less than 1/3 urn length;

exothecial cells isodiametric, walls sinuate;

stomata immersed;

annulus absent;

operculum short- to long-conic;

exostome teeth light brown to red-brown, triangular-acute;

endostome hyaline to yellow, basal membrane 1/2 exostome length or slightly longer, segments tapered apically, distinctly keeled, broadly perforate, cilia long, nodulose.

inclined 90–180°, brown to stramineous, broadly pyriform, neck 1/3 urn length;

exothecial cells short-rectangular, walls weakly to moderately sinuate;

stomata superficial;

annulus present;

operculum bluntly conic;

exostome teeth yellow to brown, triangular-acute;

endostome hyaline, basal membrane to 1/2 exostome length, segments tapered apically, distinctly keeled, broadly perforate, cilia short to long.

Spores

15–21 µm, finely roughened.

14–21 µm, finely roughened.

Specialized

asexual reproduction absent.

asexual reproduction absent.

Pohlia wahlenbergii

Pohlia ludwigii

Phenology Capsules mature spring (Apr–Jun). Capsules mature summer (Jun–Aug).
Habitat Disturbed clay or rarely sandy soil, path banks, along streams Soil, late snowmelt areas in alpine and subalpine zones
Elevation low to high elevations high elevations
Distribution
from FNA
AK; AR; AZ; CA; CO; CT; DC; IA; ID; IL; KS; KY; LA; MA; MD; ME; MI; MN; MO; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; SD; TN; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NL; NS; NT; NU; ON; PE; QC; YT; Mexico; Central America; South America; Greenland; Eurasia; Antarctica
[WildflowerSearch map]
from FNA
AK; AZ; CO; ID; MT; WA; BC; NT; ON; QC; Eurasia
Discussion

Pohlia wahlenbergii is one of the most geographically widespread and common species of the genus. The leaves are pale whitish, decurrent, and have lax, thin-walled cells. The species varies greatly in size from very slender delicate plants in suboptimal habitats to large and deep cushions in cold high altitude and latitude sites. Large plants of P. wahlenbergii typically have red stems and sometimes reddish pigmentation to the leaves as well. Some authors refer large expressions to var. glaciale (or P. glaciale), but gradation in size and pigmentation is completely continuous, and different collectors draw the line between the typical variety and var. glaciale at different points. The exothecial cell walls in capsules of this species are somewhat collenchymatous.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Pohlia ludwigii is a rare alpine species, often found in habitats similar to those of P. obtusifolia and characterized by strongly decurrent leaves that are often slightly cucullate apically. The older lower leaves generally have a pink to reddish tinge. The leaf apices are acute to bluntly acute; the endostome segments are papillose; and the perichaetial leaf costae are very strong at their bases. In late snowmelt areas, P. ludwigii can form deep cushions when sterile.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 28, p. 212. FNA vol. 28, p. 201.
Parent taxa Mielichhoferiaceae > Pohlia Mielichhoferiaceae > Pohlia
Sibling taxa
P. andalusica, P. andrewsii, P. annotina, P. atropurpurea, P. beringiensis, P. bolanderi, P. brevinervis, P. bulbifera, P. camptotrachela, P. cardotii, P. columbica, P. cruda, P. crudoides, P. drummondii, P. elongata, P. erecta, P. filum, P. flexuosa, P. lescuriana, P. longibracteata, P. longicolla, P. ludwigii, P. melanodon, P. nutans, P. obtusifolia, P. pacifica, P. proligera, P. rabunbaldensis, P. robertsonii, P. tundrae, P. vexans
P. andalusica, P. andrewsii, P. annotina, P. atropurpurea, P. beringiensis, P. bolanderi, P. brevinervis, P. bulbifera, P. camptotrachela, P. cardotii, P. columbica, P. cruda, P. crudoides, P. drummondii, P. elongata, P. erecta, P. filum, P. flexuosa, P. lescuriana, P. longibracteata, P. longicolla, P. melanodon, P. nutans, P. obtusifolia, P. pacifica, P. proligera, P. rabunbaldensis, P. robertsonii, P. tundrae, P. vexans, P. wahlenbergii
Synonyms Hypnum wahlenbergii, Mniobryum wahlenbergii Bryum ludwigii
Name authority (F. Weber & D. Mohr) A. L. Andrews: in A. J. Grout, Moss Fl. N. Amer. 2. 203. (1935) (Sprengel ex Schwagrichen) Brotherus: Acta Soc. Sci. Fenn. 19(12): 27. (1892)
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