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pale nodding-cap moss, pale pohlia, Wahlenberg's pohlia moss

pohlia moss

Habit Plants small to very large, whitish or sometimes in large forms reddish, dull. Plants medium-sized, green to yellow-green, shiny.
Stems

0.8–10 cm.

0.8–3.5 cm.

Leaves

laxly spreading to more stiffly erect, lanceolate to ovate-lanceolate, 0.6–1.2 mm;

margins weakly to strongly serrulate in distal 1/3;

costa ending well before apex;

distal medial laminal cells laxly and broadly hexagonal to rhomboidal, 65–110 µm, walls thin.

erect to ± spreading, long-lanceolate, 1.5–2.6 mm;

margins serrulate to serrate in distal 1/3;

costa subpercurrent or percurrent;

distal medial laminal cells linear- rhomboidal, vermicular, firm, 80–140 µm, walls thin.

Seta

orange-brown.

orange to orange-brown.

Sexual condition

dioicous;

perigonial leaves ovate;

perichaetial leaves weakly differentiated, lanceolate.

paroicous;

perichaetial leaves gradually differentiated, ± long-lanceolate.

Capsule

inclined ± 180°, brown to red-brown, sometimes stramineous, short-pyriform to urceolate, neck less than 1/3 urn length;

exothecial cells isodiametric, walls sinuate;

stomata immersed;

annulus absent;

operculum short- to long-conic;

exostome teeth light brown to red-brown, triangular-acute;

endostome hyaline to yellow, basal membrane 1/2 exostome length or slightly longer, segments tapered apically, distinctly keeled, broadly perforate, cilia long, nodulose.

inclined 10–90°, stramineous to orange-brown, long- and slender-pyriform, neck 1/2–1 time urn length;

exothecial cells elongate-rectangular, walls straight;

stomata superficial;

annulus present;

operculum conic;

exostome teeth yellow to brown, acute-triangular;

endostome hyaline, basal membrane 1/4–1/2 exostome length, segments narrowly keeled, narrowly to not perforate, cilia absent to rudimentary.

Spores

15–21 µm, finely roughened.

16–23 µm, distinctly roughened.

Specialized

asexual reproduction absent.

asexual reproduction absent.

Pohlia wahlenbergii

Pohlia longicolla

Phenology Capsules mature spring (Apr–Jun). Capsules mature spring–summer (Apr–Jul).
Habitat Disturbed clay or rarely sandy soil, path banks, along streams Humus-rich soil banks, along streams and paths
Elevation low to high elevations moderate to high elevations
Distribution
from FNA
AK; AR; AZ; CA; CO; CT; DC; IA; ID; IL; KS; KY; LA; MA; MD; ME; MI; MN; MO; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; SD; TN; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NL; NS; NT; NU; ON; PE; QC; YT; Mexico; Central America; South America; Greenland; Eurasia; Antarctica
[WildflowerSearch map]
from FNA
AK; MT; NC; NH; NY; OR; PA; TN; UT; VA; WY; AB; BC; NL; NT; ON; YT; Mexico; Greenland; Eurasia
Discussion

Pohlia wahlenbergii is one of the most geographically widespread and common species of the genus. The leaves are pale whitish, decurrent, and have lax, thin-walled cells. The species varies greatly in size from very slender delicate plants in suboptimal habitats to large and deep cushions in cold high altitude and latitude sites. Large plants of P. wahlenbergii typically have red stems and sometimes reddish pigmentation to the leaves as well. Some authors refer large expressions to var. glaciale (or P. glaciale), but gradation in size and pigmentation is completely continuous, and different collectors draw the line between the typical variety and var. glaciale at different points. The exothecial cell walls in capsules of this species are somewhat collenchymatous.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Pohlia longicolla, relatively rare in North America, is a handsome species characterized by relatively large, very shiny, long-lanceolate leaves. The plants are bisexual (paroicous) and commonly form sporophytes. The sporophytes have long necks, like those of P. elongata, and like that species, the endostomial segments are narrowly or not at all perforate and cilia are absent or rudimentary.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 28, p. 212. FNA vol. 28, p. 198.
Parent taxa Mielichhoferiaceae > Pohlia Mielichhoferiaceae > Pohlia
Sibling taxa
P. andalusica, P. andrewsii, P. annotina, P. atropurpurea, P. beringiensis, P. bolanderi, P. brevinervis, P. bulbifera, P. camptotrachela, P. cardotii, P. columbica, P. cruda, P. crudoides, P. drummondii, P. elongata, P. erecta, P. filum, P. flexuosa, P. lescuriana, P. longibracteata, P. longicolla, P. ludwigii, P. melanodon, P. nutans, P. obtusifolia, P. pacifica, P. proligera, P. rabunbaldensis, P. robertsonii, P. tundrae, P. vexans
P. andalusica, P. andrewsii, P. annotina, P. atropurpurea, P. beringiensis, P. bolanderi, P. brevinervis, P. bulbifera, P. camptotrachela, P. cardotii, P. columbica, P. cruda, P. crudoides, P. drummondii, P. elongata, P. erecta, P. filum, P. flexuosa, P. lescuriana, P. longibracteata, P. ludwigii, P. melanodon, P. nutans, P. obtusifolia, P. pacifica, P. proligera, P. rabunbaldensis, P. robertsonii, P. tundrae, P. vexans, P. wahlenbergii
Synonyms Hypnum wahlenbergii, Mniobryum wahlenbergii Webera longicolla
Name authority (F. Weber & D. Mohr) A. L. Andrews: in A. J. Grout, Moss Fl. N. Amer. 2. 203. (1935) (Hedwig) Lindberg: Musc. Scand., 18. (1879)
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