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pale nodding-cap moss, pale pohlia, Wahlenberg's pohlia moss

pohlia moss

Habit Plants small to very large, whitish or sometimes in large forms reddish, dull. Plants small to medium-sized, green to reddish, dull.
Stems

0.8–10 cm.

0.4–1.5 cm, green to pale orange or pink.

Leaves

laxly spreading to more stiffly erect, lanceolate to ovate-lanceolate, 0.6–1.2 mm;

margins weakly to strongly serrulate in distal 1/3;

costa ending well before apex;

distal medial laminal cells laxly and broadly hexagonal to rhomboidal, 65–110 µm, walls thin.

erect-spreading to spreading, narrowly lanceolate to lanceolate, 1.2–2.5 mm;

base not or scarcely decurrent;

margins serrate in distal 1/3;

costa subpercurrent;

distal medial laminal cells linear-hexagonal, 85–150 µm, walls thin.

Seta

orange-brown.

orange-brown.

Sexual condition

dioicous;

perigonial leaves ovate;

perichaetial leaves weakly differentiated, lanceolate.

dioicous;

perigonial leaves to 3 mm;

perichaetial leaves ± differentiated, lanceolate to linear-lanceolate.

Capsule

inclined ± 180°, brown to red-brown, sometimes stramineous, short-pyriform to urceolate, neck less than 1/3 urn length;

exothecial cells isodiametric, walls sinuate;

stomata immersed;

annulus absent;

operculum short- to long-conic;

exostome teeth light brown to red-brown, triangular-acute;

endostome hyaline to yellow, basal membrane 1/2 exostome length or slightly longer, segments tapered apically, distinctly keeled, broadly perforate, cilia long, nodulose.

inclined 145–180°, brown to stramineous, short-pyriform to urceolate, neck less than 1/3 urn length;

exothecial cells isodiametric, walls sinuate;

stomata immersed;

annulus absent;

operculum short- to long-conic;

exostome teeth yellow to yellow-brown, triangular-acute;

endostome hyaline, basal membrane 1/2 exostome length or slightly longer, segments tapered apically, distinctly keeled, broadly perforate, cilia long and nodulose, or short.

Spores

15–21 µm, finely roughened.

13–18 µm, finely roughened.

Specialized

asexual reproduction absent.

asexual reproduction absent.

Pohlia wahlenbergii

Pohlia columbica

Phenology Capsules mature spring (Apr–Jun). Capsules mature spring (May–Jul).
Habitat Disturbed clay or rarely sandy soil, path banks, along streams Disturbed soil, along streams
Elevation low to high elevations low elevations
Distribution
from FNA
AK; AR; AZ; CA; CO; CT; DC; IA; ID; IL; KS; KY; LA; MA; MD; ME; MI; MN; MO; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; SD; TN; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NL; NS; NT; NU; ON; PE; QC; YT; Mexico; Central America; South America; Greenland; Eurasia; Antarctica
[WildflowerSearch map]
from FNA
AK; WA; BC
Discussion

Pohlia wahlenbergii is one of the most geographically widespread and common species of the genus. The leaves are pale whitish, decurrent, and have lax, thin-walled cells. The species varies greatly in size from very slender delicate plants in suboptimal habitats to large and deep cushions in cold high altitude and latitude sites. Large plants of P. wahlenbergii typically have red stems and sometimes reddish pigmentation to the leaves as well. Some authors refer large expressions to var. glaciale (or P. glaciale), but gradation in size and pigmentation is completely continuous, and different collectors draw the line between the typical variety and var. glaciale at different points. The exothecial cell walls in capsules of this species are somewhat collenchymatous.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Pohlia columbica is a very rare and poorly known species characterized by dull reddish plants, variably long-acuminate perigonial leaves, pale yellow-brown exostome teeth, and immersed stomata. The exothecial cell walls of capsules of this species are somewhat collenchymatous; the perichaetial leaves are 3 mm long.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 28, p. 212. FNA vol. 28, p. 209.
Parent taxa Mielichhoferiaceae > Pohlia Mielichhoferiaceae > Pohlia
Sibling taxa
P. andalusica, P. andrewsii, P. annotina, P. atropurpurea, P. beringiensis, P. bolanderi, P. brevinervis, P. bulbifera, P. camptotrachela, P. cardotii, P. columbica, P. cruda, P. crudoides, P. drummondii, P. elongata, P. erecta, P. filum, P. flexuosa, P. lescuriana, P. longibracteata, P. longicolla, P. ludwigii, P. melanodon, P. nutans, P. obtusifolia, P. pacifica, P. proligera, P. rabunbaldensis, P. robertsonii, P. tundrae, P. vexans
P. andalusica, P. andrewsii, P. annotina, P. atropurpurea, P. beringiensis, P. bolanderi, P. brevinervis, P. bulbifera, P. camptotrachela, P. cardotii, P. cruda, P. crudoides, P. drummondii, P. elongata, P. erecta, P. filum, P. flexuosa, P. lescuriana, P. longibracteata, P. longicolla, P. ludwigii, P. melanodon, P. nutans, P. obtusifolia, P. pacifica, P. proligera, P. rabunbaldensis, P. robertsonii, P. tundrae, P. vexans, P. wahlenbergii
Synonyms Hypnum wahlenbergii, Mniobryum wahlenbergii Webera columbica
Name authority (F. Weber & D. Mohr) A. L. Andrews: in A. J. Grout, Moss Fl. N. Amer. 2. 203. (1935) (Kindberg) A. L. Andrews: in A. J. Grout, Moss Fl. N. Amer. 2: 202. (1935)
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