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pale nodding-cap moss, pale pohlia, Wahlenberg's pohlia moss

andrews' pohlia moss

Habit Plants small to very large, whitish or sometimes in large forms reddish, dull. Plants medium-sized, green, glossy.
Stems

0.8–10 cm.

0.5–2.5 cm.

Leaves

laxly spreading to more stiffly erect, lanceolate to ovate-lanceolate, 0.6–1.2 mm;

margins weakly to strongly serrulate in distal 1/3;

costa ending well before apex;

distal medial laminal cells laxly and broadly hexagonal to rhomboidal, 65–110 µm, walls thin.

± erect, lanceolate, 0.6–1.6 mm;

margins serrulate to serrate in distal 1/3;

costa subpercurrent;

distal medial laminal cells rhombic to linear-rhomboidal, 60–105 µm, walls thin.

Seta

orange-brown.

orange-brown.

Sexual condition

dioicous;

perigonial leaves ovate;

perichaetial leaves weakly differentiated, lanceolate.

dioicous;

perigonial leaves ovate;

perichaetial leaves weakly differentiated, narrowly lanceolate.

Capsule

inclined ± 180°, brown to red-brown, sometimes stramineous, short-pyriform to urceolate, neck less than 1/3 urn length;

exothecial cells isodiametric, walls sinuate;

stomata immersed;

annulus absent;

operculum short- to long-conic;

exostome teeth light brown to red-brown, triangular-acute;

endostome hyaline to yellow, basal membrane 1/2 exostome length or slightly longer, segments tapered apically, distinctly keeled, broadly perforate, cilia long, nodulose.

inclined 95–180°, brown to stramineous, pyriform, neck 1/3 urn length;

exothecial cells short-rectangular, walls sinuate;

stomata superficial;

annulus present;

operculum convex-conic;

exostome teeth yellow-brown, narrowly triangular-acute;

endostome hyaline, basal membrane 1/2 exostome length, segments distinctly keeled, broadly perforate, cilia short to rudimentary.

Spores

15–21 µm, finely roughened.

16–21 µm, finely roughened.

Specialized

asexual reproduction absent.

asexual reproduction usually present when sterile;

axillary gemmae in dense clusters, isodiametric to oblong, orange to red or sometimes green, leaf primordia restricted to apex, incurved, peglike, inconspicuous.

Pohlia wahlenbergii

Pohlia andrewsii

Phenology Capsules mature spring (Apr–Jun). Capsules mature summer (Jun–Aug).
Habitat Disturbed clay or rarely sandy soil, path banks, along streams Acid, sandy disturbed soil, path banks, stream banks
Elevation low to high elevations low to high elevations
Distribution
from FNA
AK; AR; AZ; CA; CO; CT; DC; IA; ID; IL; KS; KY; LA; MA; MD; ME; MI; MN; MO; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; SD; TN; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NL; NS; NT; NU; ON; PE; QC; YT; Mexico; Central America; South America; Greenland; Eurasia; Antarctica
[WildflowerSearch map]
from FNA
AK; NT; YT; Greenland; Europe
Discussion

Pohlia wahlenbergii is one of the most geographically widespread and common species of the genus. The leaves are pale whitish, decurrent, and have lax, thin-walled cells. The species varies greatly in size from very slender delicate plants in suboptimal habitats to large and deep cushions in cold high altitude and latitude sites. Large plants of P. wahlenbergii typically have red stems and sometimes reddish pigmentation to the leaves as well. Some authors refer large expressions to var. glaciale (or P. glaciale), but gradation in size and pigmentation is completely continuous, and different collectors draw the line between the typical variety and var. glaciale at different points. The exothecial cell walls in capsules of this species are somewhat collenchymatous.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

The glossy leaves of Pohlia andrewsii are much like those of P. proligera, but the axillary gemmae are oblong, typically orange to red, and have two to five short primordia incurved over the gemma apex.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 28, p. 212. FNA vol. 28, p. 208.
Parent taxa Mielichhoferiaceae > Pohlia Mielichhoferiaceae > Pohlia
Sibling taxa
P. andalusica, P. andrewsii, P. annotina, P. atropurpurea, P. beringiensis, P. bolanderi, P. brevinervis, P. bulbifera, P. camptotrachela, P. cardotii, P. columbica, P. cruda, P. crudoides, P. drummondii, P. elongata, P. erecta, P. filum, P. flexuosa, P. lescuriana, P. longibracteata, P. longicolla, P. ludwigii, P. melanodon, P. nutans, P. obtusifolia, P. pacifica, P. proligera, P. rabunbaldensis, P. robertsonii, P. tundrae, P. vexans
P. andalusica, P. annotina, P. atropurpurea, P. beringiensis, P. bolanderi, P. brevinervis, P. bulbifera, P. camptotrachela, P. cardotii, P. columbica, P. cruda, P. crudoides, P. drummondii, P. elongata, P. erecta, P. filum, P. flexuosa, P. lescuriana, P. longibracteata, P. longicolla, P. ludwigii, P. melanodon, P. nutans, P. obtusifolia, P. pacifica, P. proligera, P. rabunbaldensis, P. robertsonii, P. tundrae, P. vexans, P. wahlenbergii
Synonyms Hypnum wahlenbergii, Mniobryum wahlenbergii
Name authority (F. Weber & D. Mohr) A. L. Andrews: in A. J. Grout, Moss Fl. N. Amer. 2. 203. (1935) A. J. Shaw: Bryologist 84: 70, figs. 11 – 34. (1981)
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