Pohlia nutans
Pohlia beringiensis
nodding pohlia, nodding thread-moss, pohlia moss
Bering's pohlia moss
0.5–3.5(–10) cm.
0.5–2 cm.
erect to ± spreading, lanceolate to ovate-lanceolate, 2 mm;
base not or scarcely decurrent;
margins subentire or more often serrulate to serrate in distal 1/3;
costa subpercurrent, percurrent, or rarely short-excurrent;
distal medial laminal cells hexagonal to rhomboidal, firm, 50–90 µm, walls moderately thick.
± erect, unaltered when dry, lanceolate, 0.9–1.5 mm;
margins serrulate to serrate in distal 1/3;
costa subpercurrent;
distal medial laminal cells linear-rhomboidal, vermicular, 60–100 µm, walls thin.
orange to orange-brown.
paroicous, rarely dioicous;
perigonial leaves ovate-lanceolate;
perichaetial leaves somewhat differentiated, ± long-lanceolate.
unknown, presumed dioicous.
inclined 80–100°, orange to orange-brown, slender-pyriform, neck 1/2 urn length;
exothecial cells elongate-rectangular, walls straight;
stomata superficial;
annulus present;
operculum conic;
exostome teeth yellow to orange-brown, acute-triangular;
endostome hyaline or rarely orange-brown, basal membrane 1/2 exostome length, segments tapered apically, broadly keeled, broadly perforate, cilia short to long.
16–22 µm, finely to distinctly roughened.
asexual reproduction typically absent.
asexual reproduction usually present;
axillary gemmae single, bulbiform, red, darker to black when dry, leaf primordia at apex and more proximally, laminate, large, flexuose.
unknown.
Pohlia nutans
Pohlia beringiensis
Pohlia nutans is the most common species of Pohlia in North America, Europe, and elsewhere in the Northern Hemisphere. In contrast to the treatment by A. J. Shaw (1982), P. sphagnicola is not separated here from P. nutans; the type of P. sphagnicola is European. Plants referable to P. sphagnicola, including North American collections, differ in being dioicous and having entire leaves, slightly shorter laminal cells, and smaller spores. The habitat in Sphagnum hummocks is not diagnostic; at least 80% of plants growing in Sphagnum are P. nutans. Pohlia schimperi, a northern form with reddish leaves, also dioicous, is not recognized, as per Shaw.
The peristome of Pohlia nutans is as well developed as any in Pohlia, with long, tapered, trabeculate exostome teeth and well-developed endostomes with broadly keeled, widely perforate segments and short to long cilia. The laminal cells are short- to elongate-hexagonal with thickened walls. The only other North American species with relatively thick-walled laminal cells is P. elongata; the cells of P. elongata are typically longer, but there is extensive overlap in sizes. Plants without sporophytes have generally been named P. nutans, although some collections from montane regions may actually be P. elongata; sporophytes of P. elongata have longer necks, as long as or longer than the urns, and reduced peristomes with endostome segments narrowly split along the keel and (usually) no cilia.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Pohlia beringiensis has cherry red stems with erect, whitish leaves and red bulbiform axillary gemmae. The gemmae are much like those of P. drummondii, but are even more red and contrast conspicuously with the whitish leaves. The organic-poor substrates on which P. beringiensis occurs are quite different from the relatively humic soils on which P. drummondii occurs.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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