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opal pohlia, opal thread-moss, pohlia moss

Habit Plants medium-sized to large, green, whitish, or sometimes bluish tinged, shiny. Plants small to medium-sized, green, dull.
Stems

0.5–2.5 cm.

0.5–1 cm.

Leaves

erect to ± spreading, lanceolate to elliptic, 0.8–2 mm;

margins serrulate to serrate in distal 1/3;

costa subpercurrent;

distal medial laminal cells linear-rhomboidal, vermicular, firm, 70–140 µm, walls thin.

erect to ± spreading, lanceolate, 1.3–1.6 mm;

margins serrulate to serrate in distal 1/3;

costa subpercurrent;

distal medial laminal cells irregularly hexagonal to rhomboidal, 70–120 µm, walls thin to firm.

Seta

orange to orange-brown.

multiple, orange-brown.

Sexual condition

paroicous, rarely dioicous;

perigonial leaves in dioicous plants linear- or long-lanceolate from ovate base;

perichaetial leaves strongly or rarely weakly differentiated, linear-lanceolate.

paroicous;

perichaetial leaves scarcely differentiated, lanceolate.

Capsule

inclined 10–135°, stramineous to orange-brown, long- and slender-pyriform, neck 1/2 urn length;

exothecial cells elongate-rectangular, walls straight;

stomata superficial;

annulus present;

operculum conic;

exostome teeth yellow-brown to red-brown, acute-triangular;

endostome hyaline, basal membrane 1/2 exostome length, segments broadly keeled, broadly perforate, cilia short to long.

inclined 10–45°, stramineous to orange-brown, long- and slender-pyriform, neck 3/4–1 time urn length;

exothecial cells elongate-rectangular, walls slightly sinuate;

stomata superficial;

annulus present;

operculum long-conic;

exostome teeth orange-yellow to brown, bluntly acute;

endostome hyaline to yellow-brown, basal membrane low, scarcely exceeding capsule rim, segments narrow, keeled basally, not or slightly perforate, cilia absent or rudimentary.

Spores

18–26 µm, ± coarsely papillose.

16–20 µm, ± finely but distinctly roughened.

Specialized

asexual reproduction absent.

asexual reproduction absent.

Pohlia cruda

Pohlia robertsonii

Phenology Capsules mature summer (Jun–Aug). Capsules mature spring (Mar–Apr).
Habitat Soil banks, rock crevices, under roots, tundra soil and paths Sandy or gravelly soil, base of sandstone or volcanic rock in open deciduous woodlands
Elevation moderate to high elevations low elevations
Distribution
from FNA
AK; AZ; CA; CO; IA; ID; MA; MD; ME; MI; MN; MT; NC; NE; NJ; NM; NV; NY; OR; PA; SD; TN; UT; VA; VT; WA; WV; WY; AB; BC; MB; NB; NL; NS; NT; NU; ON; QC; SK; YT; Mexico; South America; Australia; Greenland; Pacific Islands; Eurasia; Antarctica
[WildflowerSearch map]
from FNA
CA
Discussion

Pohlia cruda is distinguished by the glossy, pale green to whitish or bluish leaves that are narrowly elliptic to lanceolate; the laminal cells are long and narrowly linear-vermicular; the perichaetial leaves are strongly differentiated and linear-lanceolate; the sporophytes have necks about as long as the urn, with long-tapered exostome teeth, broadly keeled and perforate segments, and short to long cilia. Along with P. nutans, P. cruda is one of the most common North American species of the genus. Unlike P. nutans, P. cruda is restricted to northern and montane sites where it grows in rock crevices and on soil banks. The leaves of this species are sometimes slightly complanate. The gametophytes can be unisexual (dioicous) or bisexual (paroicous).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Pohlia robertsonii, known only from Monterey County, is the only North American polysetous species of Pohlia. The leaves are less dull in appearance than those of P. elongata and P. nutans, but not notably shiny like those of P. cruda or P. longicolla; they are more like the moderately glossy leaves often seen in P. bolanderi. The laminal cells have firm but not thick walls, thinner-walled than in P. elongata and P. nutans, shorter and less vermicular than in P. cruda or P. longicolla. Sporophytes are produced abundantly on the paroicous gametophytes. The exostome teeth are rather irregular and poorly developed, not gradually tapered to narrow acute apices, and the endostome segments are narrow, scarcely keeled, and narrowly or not at all perforate.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 28, p. 198. FNA vol. 28, p. 199.
Parent taxa Mielichhoferiaceae > Pohlia Mielichhoferiaceae > Pohlia
Sibling taxa
P. andalusica, P. andrewsii, P. annotina, P. atropurpurea, P. beringiensis, P. bolanderi, P. brevinervis, P. bulbifera, P. camptotrachela, P. cardotii, P. columbica, P. crudoides, P. drummondii, P. elongata, P. erecta, P. filum, P. flexuosa, P. lescuriana, P. longibracteata, P. longicolla, P. ludwigii, P. melanodon, P. nutans, P. obtusifolia, P. pacifica, P. proligera, P. rabunbaldensis, P. robertsonii, P. tundrae, P. vexans, P. wahlenbergii
P. andalusica, P. andrewsii, P. annotina, P. atropurpurea, P. beringiensis, P. bolanderi, P. brevinervis, P. bulbifera, P. camptotrachela, P. cardotii, P. columbica, P. cruda, P. crudoides, P. drummondii, P. elongata, P. erecta, P. filum, P. flexuosa, P. lescuriana, P. longibracteata, P. longicolla, P. ludwigii, P. melanodon, P. nutans, P. obtusifolia, P. pacifica, P. proligera, P. rabunbaldensis, P. tundrae, P. vexans, P. wahlenbergii
Synonyms Mnium crudum
Name authority (Hedwig) Lindberg: Musc. Scand., 18. (1879) Shevock & A. J. Shaw: Bryologist 108: 177, fig. 1. (2005)
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