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opal pohlia, opal thread-moss, pohlia moss

ludwig's pohlia moss, ludwig's thread-moss

Habit Plants medium-sized to large, green, whitish, or sometimes bluish tinged, shiny. Plants medium-sized to large, green, sometimes red tinged, dull.
Stems

0.5–2.5 cm.

0.8–3.5 cm.

Leaves

erect to ± spreading, lanceolate to elliptic, 0.8–2 mm;

margins serrulate to serrate in distal 1/3;

costa subpercurrent;

distal medial laminal cells linear-rhomboidal, vermicular, firm, 70–140 µm, walls thin.

erect to ± spreading, lanceolate to broadly lanceolate, 1.2–2.4 mm;

base long-decurrent;

margins serrulate to serrate in distal 1/3;

costa subpercurrent;

distal medial laminal cells rhombic to rhomboidal, 40–80 µm, walls firm to thin.

Seta

orange to orange-brown.

orange-brown.

Sexual condition

paroicous, rarely dioicous;

perigonial leaves in dioicous plants linear- or long-lanceolate from ovate base;

perichaetial leaves strongly or rarely weakly differentiated, linear-lanceolate.

dioicous;

perigonial leaves broadly short-ovate;

perichaetial leaves scarcely differentiated, broadly lanceolate.

Capsule

inclined 10–135°, stramineous to orange-brown, long- and slender-pyriform, neck 1/2 urn length;

exothecial cells elongate-rectangular, walls straight;

stomata superficial;

annulus present;

operculum conic;

exostome teeth yellow-brown to red-brown, acute-triangular;

endostome hyaline, basal membrane 1/2 exostome length, segments broadly keeled, broadly perforate, cilia short to long.

inclined 90–180°, brown to stramineous, broadly pyriform, neck 1/3 urn length;

exothecial cells short-rectangular, walls weakly to moderately sinuate;

stomata superficial;

annulus present;

operculum bluntly conic;

exostome teeth yellow to brown, triangular-acute;

endostome hyaline, basal membrane to 1/2 exostome length, segments tapered apically, distinctly keeled, broadly perforate, cilia short to long.

Spores

18–26 µm, ± coarsely papillose.

14–21 µm, finely roughened.

Specialized

asexual reproduction absent.

asexual reproduction absent.

Pohlia cruda

Pohlia ludwigii

Phenology Capsules mature summer (Jun–Aug). Capsules mature summer (Jun–Aug).
Habitat Soil banks, rock crevices, under roots, tundra soil and paths Soil, late snowmelt areas in alpine and subalpine zones
Elevation moderate to high elevations high elevations
Distribution
from FNA
AK; AZ; CA; CO; IA; ID; MA; MD; ME; MI; MN; MT; NC; NE; NJ; NM; NV; NY; OR; PA; SD; TN; UT; VA; VT; WA; WV; WY; AB; BC; MB; NB; NL; NS; NT; NU; ON; QC; SK; YT; Mexico; South America; Australia; Greenland; Pacific Islands; Eurasia; Antarctica
[WildflowerSearch map]
from FNA
AK; AZ; CO; ID; MT; WA; BC; NT; ON; QC; Eurasia
Discussion

Pohlia cruda is distinguished by the glossy, pale green to whitish or bluish leaves that are narrowly elliptic to lanceolate; the laminal cells are long and narrowly linear-vermicular; the perichaetial leaves are strongly differentiated and linear-lanceolate; the sporophytes have necks about as long as the urn, with long-tapered exostome teeth, broadly keeled and perforate segments, and short to long cilia. Along with P. nutans, P. cruda is one of the most common North American species of the genus. Unlike P. nutans, P. cruda is restricted to northern and montane sites where it grows in rock crevices and on soil banks. The leaves of this species are sometimes slightly complanate. The gametophytes can be unisexual (dioicous) or bisexual (paroicous).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Pohlia ludwigii is a rare alpine species, often found in habitats similar to those of P. obtusifolia and characterized by strongly decurrent leaves that are often slightly cucullate apically. The older lower leaves generally have a pink to reddish tinge. The leaf apices are acute to bluntly acute; the endostome segments are papillose; and the perichaetial leaf costae are very strong at their bases. In late snowmelt areas, P. ludwigii can form deep cushions when sterile.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 28, p. 198. FNA vol. 28, p. 201.
Parent taxa Mielichhoferiaceae > Pohlia Mielichhoferiaceae > Pohlia
Sibling taxa
P. andalusica, P. andrewsii, P. annotina, P. atropurpurea, P. beringiensis, P. bolanderi, P. brevinervis, P. bulbifera, P. camptotrachela, P. cardotii, P. columbica, P. crudoides, P. drummondii, P. elongata, P. erecta, P. filum, P. flexuosa, P. lescuriana, P. longibracteata, P. longicolla, P. ludwigii, P. melanodon, P. nutans, P. obtusifolia, P. pacifica, P. proligera, P. rabunbaldensis, P. robertsonii, P. tundrae, P. vexans, P. wahlenbergii
P. andalusica, P. andrewsii, P. annotina, P. atropurpurea, P. beringiensis, P. bolanderi, P. brevinervis, P. bulbifera, P. camptotrachela, P. cardotii, P. columbica, P. cruda, P. crudoides, P. drummondii, P. elongata, P. erecta, P. filum, P. flexuosa, P. lescuriana, P. longibracteata, P. longicolla, P. melanodon, P. nutans, P. obtusifolia, P. pacifica, P. proligera, P. rabunbaldensis, P. robertsonii, P. tundrae, P. vexans, P. wahlenbergii
Synonyms Mnium crudum Bryum ludwigii
Name authority (Hedwig) Lindberg: Musc. Scand., 18. (1879) (Sprengel ex Schwagrichen) Brotherus: Acta Soc. Sci. Fenn. 19(12): 27. (1892)
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