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opal pohlia, opal thread-moss, pohlia moss

Habit Plants medium-sized to large, green, whitish, or sometimes bluish tinged, shiny. Plants small, green, dull.
Stems

0.5–2.5 cm.

0.5–1.5 cm.

Leaves

erect to ± spreading, lanceolate to elliptic, 0.8–2 mm;

margins serrulate to serrate in distal 1/3;

costa subpercurrent;

distal medial laminal cells linear-rhomboidal, vermicular, firm, 70–140 µm, walls thin.

± erect, lanceolate, 0.6–1.3 mm;

margins serrulate to serrate in distal 1/3;

costa subpercurrent;

distal medial laminal cells rhombic to linear-rhomboidal, 60–105 µm, walls thin.

Seta

orange to orange-brown.

orange-brown.

Sexual condition

paroicous, rarely dioicous;

perigonial leaves in dioicous plants linear- or long-lanceolate from ovate base;

perichaetial leaves strongly or rarely weakly differentiated, linear-lanceolate.

Capsule

inclined 10–135°, stramineous to orange-brown, long- and slender-pyriform, neck 1/2 urn length;

exothecial cells elongate-rectangular, walls straight;

stomata superficial;

annulus present;

operculum conic;

exostome teeth yellow-brown to red-brown, acute-triangular;

endostome hyaline, basal membrane 1/2 exostome length, segments broadly keeled, broadly perforate, cilia short to long.

inclined 95–180°, brown to stramineous, pyriform, neck 1/3 urn length;

exothecial cells short-rectangular, walls sinuate;

stomata superficial;annulus present;

operculum convex-conic;

exostome teeth yellow-brown, narrowly triangular-acute;

endostome hyaline, basal membrane 1/2 exostome length, segments distinctly keeled, broadly perforate, cilia short to rudimentary.

Spores

18–26 µm, ± coarsely papillose.

16–21 µm, finely roughened].

Specialized

asexual reproduction absent.

asexual reproduction usually present when sterile;

axillary gemmae in dense clusters, dimorphic, isodiametric to oblong, knobby in outline and brown, or narrowly linear-vermicular and hyaline, leaf primordia absent or 1 or 2, inconspicuous, restricted to apex, peglike.

[sexual condition dioicous; perigonial leaves ovate; perichaetial leaves weakly differentiated, narrowly lanceolate.

Pohlia cruda

Pohlia flexuosa

Phenology Capsules mature summer (Jun–Aug). Capsule maturity unknown.
Habitat Soil banks, rock crevices, under roots, tundra soil and paths Acid, sandy disturbed soil, path banks, stream banks
Elevation moderate to high elevations low elevations
Distribution
from FNA
AK; AZ; CA; CO; IA; ID; MA; MD; ME; MI; MN; MT; NC; NE; NJ; NM; NV; NY; OR; PA; SD; TN; UT; VA; VT; WA; WV; WY; AB; BC; MB; NB; NL; NS; NT; NU; ON; QC; SK; YT; Mexico; South America; Australia; Greenland; Pacific Islands; Eurasia; Antarctica
[WildflowerSearch map]
from FNA
CA; Europe; Asia
Discussion

Pohlia cruda is distinguished by the glossy, pale green to whitish or bluish leaves that are narrowly elliptic to lanceolate; the laminal cells are long and narrowly linear-vermicular; the perichaetial leaves are strongly differentiated and linear-lanceolate; the sporophytes have necks about as long as the urn, with long-tapered exostome teeth, broadly keeled and perforate segments, and short to long cilia. Along with P. nutans, P. cruda is one of the most common North American species of the genus. Unlike P. nutans, P. cruda is restricted to northern and montane sites where it grows in rock crevices and on soil banks. The leaves of this species are sometimes slightly complanate. The gametophytes can be unisexual (dioicous) or bisexual (paroicous).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

The slender, dull plants of Pohlia flexuosa, which have leaves somewhat contorted when dry, are like those of P. annotina and P. camptotrachela. The gemmae are dimorphic, and either hyaline to whitish and narrowly linear-vermicular, with one or two poorly differentiated leaf primordia, or small, blackish, and oblong to globose, with angular cells. This species is known from only one North American site in Yuba County but is common in parts of Asia including China, and it occurs in western Europe.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 28, p. 198. FNA vol. 28, p. 208.
Parent taxa Mielichhoferiaceae > Pohlia Mielichhoferiaceae > Pohlia
Sibling taxa
P. andalusica, P. andrewsii, P. annotina, P. atropurpurea, P. beringiensis, P. bolanderi, P. brevinervis, P. bulbifera, P. camptotrachela, P. cardotii, P. columbica, P. crudoides, P. drummondii, P. elongata, P. erecta, P. filum, P. flexuosa, P. lescuriana, P. longibracteata, P. longicolla, P. ludwigii, P. melanodon, P. nutans, P. obtusifolia, P. pacifica, P. proligera, P. rabunbaldensis, P. robertsonii, P. tundrae, P. vexans, P. wahlenbergii
P. andalusica, P. andrewsii, P. annotina, P. atropurpurea, P. beringiensis, P. bolanderi, P. brevinervis, P. bulbifera, P. camptotrachela, P. cardotii, P. columbica, P. cruda, P. crudoides, P. drummondii, P. elongata, P. erecta, P. filum, P. lescuriana, P. longibracteata, P. longicolla, P. ludwigii, P. melanodon, P. nutans, P. obtusifolia, P. pacifica, P. proligera, P. rabunbaldensis, P. robertsonii, P. tundrae, P. vexans, P. wahlenbergii
Synonyms Mnium crudum
Name authority (Hedwig) Lindberg: Musc. Scand., 18. (1879) Hooker: Icon. Pl. 1: plate 19, fig. 5. (1836)
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