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opal pohlia, opal thread-moss, pohlia moss

elongate pohlia moss, long-fruit pohlia, long-fruit thread-moss

Habit Plants medium-sized to large, green, whitish, or sometimes bluish tinged, shiny. Plants small, green, dull.
Stems

0.5–2.5 cm.

0.5–2.5 cm.

Leaves

erect to ± spreading, lanceolate to elliptic, 0.8–2 mm;

margins serrulate to serrate in distal 1/3;

costa subpercurrent;

distal medial laminal cells linear-rhomboidal, vermicular, firm, 70–140 µm, walls thin.

erect to ± spreading, lanceolate to ovate-lanceolate, 2 mm;

margins subentire or more often serrulate to serrate in distal 1/3;

costa subpercurrent or percurrent;

distal medial laminal cells hexagonal to rhomboidal, firm, 45–85 µm, walls moderately thick.

Seta

orange to orange-brown.

orange to orange-brown.

Sexual condition

paroicous, rarely dioicous;

perigonial leaves in dioicous plants linear- or long-lanceolate from ovate base;

perichaetial leaves strongly or rarely weakly differentiated, linear-lanceolate.

paroicous, autoicous, or rarely dioicous;

perigonial leaves in dioicous plants broadly short-ovate;

perichaetial leaves somewhat differentiated, ± long-lanceolate.

Capsule

inclined 10–135°, stramineous to orange-brown, long- and slender-pyriform, neck 1/2 urn length;

exothecial cells elongate-rectangular, walls straight;

stomata superficial;

annulus present;

operculum conic;

exostome teeth yellow-brown to red-brown, acute-triangular;

endostome hyaline, basal membrane 1/2 exostome length, segments broadly keeled, broadly perforate, cilia short to long.

inclined 10–90°, stramineous, orange, or orange-brown, short- to long- and slender-pyriform, neck 1/2–1 time urn length;

exothecial cells elongate-rectangular, walls straight;

stomata superficial;

annulus present;

operculum conic;

exostome teeth yellow to brown, acute-triangular;

endostome hyaline, basal membrane barely exceeding capsule rim or to 1/3 exostome length, segments narrowly or not keeled, narrowly to not perforate, cilia absent to rudimentary, rarely long.

Spores

18–26 µm, ± coarsely papillose.

16–23 µm, distinctly roughened.

Specialized

asexual reproduction absent.

asexual reproduction absent.

Pohlia cruda

Pohlia elongata

Phenology Capsules mature summer (Jun–Aug). Capsules mature spring–summer (Apr–Jul).
Habitat Soil banks, rock crevices, under roots, tundra soil and paths Humus-rich soil banks, along streams and paths, tree bases
Elevation moderate to high elevations moderate to high elevations
Distribution
from FNA
AK; AZ; CA; CO; IA; ID; MA; MD; ME; MI; MN; MT; NC; NE; NJ; NM; NV; NY; OR; PA; SD; TN; UT; VA; VT; WA; WV; WY; AB; BC; MB; NB; NL; NS; NT; NU; ON; QC; SK; YT; Mexico; South America; Australia; Greenland; Pacific Islands; Eurasia; Antarctica
[WildflowerSearch map]
from FNA
AK; AL; AZ; CA; CO; GA; IA; ME; MI; MN; MT; NC; NH; NM; NY; OH; TN; UT; VA; VT; WA; WI; BC; NB; NL; NS; NT; ON; QC; YT; Mexico; Central America; South America; Eurasia; Africa; Pacific Islands; Australia
Discussion

Pohlia cruda is distinguished by the glossy, pale green to whitish or bluish leaves that are narrowly elliptic to lanceolate; the laminal cells are long and narrowly linear-vermicular; the perichaetial leaves are strongly differentiated and linear-lanceolate; the sporophytes have necks about as long as the urn, with long-tapered exostome teeth, broadly keeled and perforate segments, and short to long cilia. Along with P. nutans, P. cruda is one of the most common North American species of the genus. Unlike P. nutans, P. cruda is restricted to northern and montane sites where it grows in rock crevices and on soil banks. The leaves of this species are sometimes slightly complanate. The gametophytes can be unisexual (dioicous) or bisexual (paroicous).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Pohlia elongata is widespread in North America but not common. Plants generally occur scattered or in small patches and regularly produce sporophytes from bisexual (paroicous) gametophytes. The capsule necks of P. elongata are typically longer than the urn and the endostome is reduced (see under 1. P. nutans). The endostomial segments vary from relatively broad and distinctly keeled with narrow perforations to narrowly linear, scarcely keeled, and not at all split along the keels. A high elevation expression that has generally been collected from exposed sites and often called P. acuminata or P. polymorpha, is characterized by small plants with relatively broad leaves and short, barrel-shaped sporophytes with short necks.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 28, p. 198. FNA vol. 28, p. 197.
Parent taxa Mielichhoferiaceae > Pohlia Mielichhoferiaceae > Pohlia
Sibling taxa
P. andalusica, P. andrewsii, P. annotina, P. atropurpurea, P. beringiensis, P. bolanderi, P. brevinervis, P. bulbifera, P. camptotrachela, P. cardotii, P. columbica, P. crudoides, P. drummondii, P. elongata, P. erecta, P. filum, P. flexuosa, P. lescuriana, P. longibracteata, P. longicolla, P. ludwigii, P. melanodon, P. nutans, P. obtusifolia, P. pacifica, P. proligera, P. rabunbaldensis, P. robertsonii, P. tundrae, P. vexans, P. wahlenbergii
P. andalusica, P. andrewsii, P. annotina, P. atropurpurea, P. beringiensis, P. bolanderi, P. brevinervis, P. bulbifera, P. camptotrachela, P. cardotii, P. columbica, P. cruda, P. crudoides, P. drummondii, P. erecta, P. filum, P. flexuosa, P. lescuriana, P. longibracteata, P. longicolla, P. ludwigii, P. melanodon, P. nutans, P. obtusifolia, P. pacifica, P. proligera, P. rabunbaldensis, P. robertsonii, P. tundrae, P. vexans, P. wahlenbergii
Synonyms Mnium crudum P. acuminata, P. elongata var. greenii, P. polymorpha
Name authority (Hedwig) Lindberg: Musc. Scand., 18. (1879) Hedwig: Sp. Musc. Frond., 171. (1801)
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