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opal pohlia, opal thread-moss, pohlia moss

Drummond's pohlia moss, Drummond's thread-moss

Habit Plants medium-sized to large, green, whitish, or sometimes bluish tinged, shiny. Plants medium-sized, dark green to reddish, glossy.
Stems

0.5–2.5 cm.

0.5–3.5 cm.

Leaves

erect to ± spreading, lanceolate to elliptic, 0.8–2 mm;

margins serrulate to serrate in distal 1/3;

costa subpercurrent;

distal medial laminal cells linear-rhomboidal, vermicular, firm, 70–140 µm, walls thin.

± spreading, unaltered when dry, lanceolate to ovate-lanceolate, 0.9–1.5 mm;

margins serrulate to serrate in distal 1/3;

costa subpercurrent;

distal medial laminal cells rhombic to rhomboidal, 50–95 µm, walls thin.

Seta

orange to orange-brown.

orange-brown.

Sexual condition

paroicous, rarely dioicous;

perigonial leaves in dioicous plants linear- or long-lanceolate from ovate base;

perichaetial leaves strongly or rarely weakly differentiated, linear-lanceolate.

dioicous;

perigonial leaves ovate;

perichaetial leaves scarcely differentiated, ovate-lanceolate.

Capsule

inclined 10–135°, stramineous to orange-brown, long- and slender-pyriform, neck 1/2 urn length;

exothecial cells elongate-rectangular, walls straight;

stomata superficial;

annulus present;

operculum conic;

exostome teeth yellow-brown to red-brown, acute-triangular;

endostome hyaline, basal membrane 1/2 exostome length, segments broadly keeled, broadly perforate, cilia short to long.

inclined 95–180°, brown to stramineous, pyriform, neck 1/3 urn length;

exothecial cells short-rectangular, walls sinuate;

stomata superficial;

annulus present;

operculum convex-conic;

exostome teeth yellow-brown, narrowly triangular-acute;

endostome hyaline, basal membrane 1/2 exostome length, segments distinctly keeled, broadly perforate, cilia short to rudimentary.

Spores

18–26 µm, ± coarsely papillose.

16–21 µm, finely roughened.

Specialized

asexual reproduction absent.

asexual reproduction usually present when sterile;

axillary gemmae single, bulbiform, cherry red, ± black when dry, leaf primordia at gemma apex and below, laminate, large, flexuose.

Pohlia cruda

Pohlia drummondii

Phenology Capsules mature summer (Jun–Aug). Capsules mature summer (Jun–Aug).
Habitat Soil banks, rock crevices, under roots, tundra soil and paths Acid, relatively humus-rich soil, alpine tundra, stream banks, path banks
Elevation moderate to high elevations low to high elevations
Distribution
from FNA
AK; AZ; CA; CO; IA; ID; MA; MD; ME; MI; MN; MT; NC; NE; NJ; NM; NV; NY; OR; PA; SD; TN; UT; VA; VT; WA; WV; WY; AB; BC; MB; NB; NL; NS; NT; NU; ON; QC; SK; YT; Mexico; South America; Australia; Greenland; Pacific Islands; Eurasia; Antarctica
[WildflowerSearch map]
from FNA
AK; AZ; CA; CO; ID; MN; MT; NH; NM; NV; OR; UT; VT; WA; WY; AB; BC; NL; NT; NU; ON; QC; YT; Greenland; Europe
Discussion

Pohlia cruda is distinguished by the glossy, pale green to whitish or bluish leaves that are narrowly elliptic to lanceolate; the laminal cells are long and narrowly linear-vermicular; the perichaetial leaves are strongly differentiated and linear-lanceolate; the sporophytes have necks about as long as the urn, with long-tapered exostome teeth, broadly keeled and perforate segments, and short to long cilia. Along with P. nutans, P. cruda is one of the most common North American species of the genus. Unlike P. nutans, P. cruda is restricted to northern and montane sites where it grows in rock crevices and on soil banks. The leaves of this species are sometimes slightly complanate. The gametophytes can be unisexual (dioicous) or bisexual (paroicous).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Pohlia drummondii has red stems (when moist) with cherry red (when fresh) bulbiform gemmae in the leaf axils. The gemmae look like small branches, with flexuose, laminate leaf primordia, and occur singly or occasionally in pairs in one to five distal leaf axils. The leaves of P. drummondii are carinate and can be similar to those of P. obtusifolia, but are not cucullate, and the plants are dioicous.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 28, p. 198. FNA vol. 28, p. 203.
Parent taxa Mielichhoferiaceae > Pohlia Mielichhoferiaceae > Pohlia
Sibling taxa
P. andalusica, P. andrewsii, P. annotina, P. atropurpurea, P. beringiensis, P. bolanderi, P. brevinervis, P. bulbifera, P. camptotrachela, P. cardotii, P. columbica, P. crudoides, P. drummondii, P. elongata, P. erecta, P. filum, P. flexuosa, P. lescuriana, P. longibracteata, P. longicolla, P. ludwigii, P. melanodon, P. nutans, P. obtusifolia, P. pacifica, P. proligera, P. rabunbaldensis, P. robertsonii, P. tundrae, P. vexans, P. wahlenbergii
P. andalusica, P. andrewsii, P. annotina, P. atropurpurea, P. beringiensis, P. bolanderi, P. brevinervis, P. bulbifera, P. camptotrachela, P. cardotii, P. columbica, P. cruda, P. crudoides, P. elongata, P. erecta, P. filum, P. flexuosa, P. lescuriana, P. longibracteata, P. longicolla, P. ludwigii, P. melanodon, P. nutans, P. obtusifolia, P. pacifica, P. proligera, P. rabunbaldensis, P. robertsonii, P. tundrae, P. vexans, P. wahlenbergii
Synonyms Mnium crudum Bryum drummondii, P. commutata
Name authority (Hedwig) Lindberg: Musc. Scand., 18. (1879) (Müller Hal.) A. L. Andrews: in A. J. Grout, Moss Fl. N. Amer. 2: 196. (1935)
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