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opal pohlia, opal thread-moss, pohlia moss

pohlia moss

Habit Plants medium-sized to large, green, whitish, or sometimes bluish tinged, shiny. Plants medium-sized to large, green, dull.
Stems

0.5–2.5 cm.

0.4–2.5 cm.

Leaves

erect to ± spreading, lanceolate to elliptic, 0.8–2 mm;

margins serrulate to serrate in distal 1/3;

costa subpercurrent;

distal medial laminal cells linear-rhomboidal, vermicular, firm, 70–140 µm, walls thin.

erect to ± spreading, narrowly long-lanceolate, 1–2.2 mm;

margins strongly revolute, serrulate to serrate in distal 1/3;

costa subpercurrent;

distal medial laminal cells linear-hexagonal, 35–80 µm, walls thin, not porose.

Seta

orange to orange-brown.

orange-brown.

Sexual condition

paroicous, rarely dioicous;

perigonial leaves in dioicous plants linear- or long-lanceolate from ovate base;

perichaetial leaves strongly or rarely weakly differentiated, linear-lanceolate.

dioicous;

perigonial leaves broadly short-ovate;

perichaetial leaves moderately differentiated, narrowly lanceolate.

Capsule

inclined 10–135°, stramineous to orange-brown, long- and slender-pyriform, neck 1/2 urn length;

exothecial cells elongate-rectangular, walls straight;

stomata superficial;

annulus present;

operculum conic;

exostome teeth yellow-brown to red-brown, acute-triangular;

endostome hyaline, basal membrane 1/2 exostome length, segments broadly keeled, broadly perforate, cilia short to long.

inclined 30–90°, brown, ovoid-cylindric, neck 1/3 urn length;

exothecial cells short- to long-rectangular, walls straight or weakly sinuate;

stomata superficial;

annulus present;

operculum conic;

exostome teeth whitish to yellow, bluntly acute, coarsely papillose throughout;

endostome whitish to hyaline, basal membrane low, to 1/3 exostome length, segments narrow, scarcely keeled, narrowly or not perforate, cilia absent or rarely rudimentary.

Spores

18–26 µm, ± coarsely papillose.

15–25 µm, finely roughened.

Specialized

asexual reproduction absent.

asexual reproduction absent.

Pohlia cruda

Pohlia crudoides

Phenology Capsules mature summer (Jun–Aug). Capsules mature summer (Jun–Aug).
Habitat Soil banks, rock crevices, under roots, tundra soil and paths Soil, tundra, on banks and in depressions
Elevation moderate to high elevations low to high elevations
Distribution
from FNA
AK; AZ; CA; CO; IA; ID; MA; MD; ME; MI; MN; MT; NC; NE; NJ; NM; NV; NY; OR; PA; SD; TN; UT; VA; VT; WA; WV; WY; AB; BC; MB; NB; NL; NS; NT; NU; ON; QC; SK; YT; Mexico; South America; Australia; Greenland; Pacific Islands; Eurasia; Antarctica
[WildflowerSearch map]
from FNA
AK; AB; BC; NT; NU; QC; YT; Greenland; Eurasia
Discussion

Pohlia cruda is distinguished by the glossy, pale green to whitish or bluish leaves that are narrowly elliptic to lanceolate; the laminal cells are long and narrowly linear-vermicular; the perichaetial leaves are strongly differentiated and linear-lanceolate; the sporophytes have necks about as long as the urn, with long-tapered exostome teeth, broadly keeled and perforate segments, and short to long cilia. Along with P. nutans, P. cruda is one of the most common North American species of the genus. Unlike P. nutans, P. cruda is restricted to northern and montane sites where it grows in rock crevices and on soil banks. The leaves of this species are sometimes slightly complanate. The gametophytes can be unisexual (dioicous) or bisexual (paroicous).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Pohlia crudoides is a distinctive Arctic species characterized by cherry red stems and relatively stiff, dark green leaves with strongly recurved margins. The perichaetial leaves of most Pohlia species have recurved margins, but the only other North American species with strongly recurved vegetative leaf margins is P. cardotii. The latter has shorter, broader leaves with short-rhombic cells often with porose-thickened walls. Pohlia crudoides has weakly tapered exostome teeth coarsely papillose from top to bottom, which give them a whitish appearance under a dissecting microscope; the endostome segments also are papillose.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 28, p. 198. FNA vol. 28, p. 200.
Parent taxa Mielichhoferiaceae > Pohlia Mielichhoferiaceae > Pohlia
Sibling taxa
P. andalusica, P. andrewsii, P. annotina, P. atropurpurea, P. beringiensis, P. bolanderi, P. brevinervis, P. bulbifera, P. camptotrachela, P. cardotii, P. columbica, P. crudoides, P. drummondii, P. elongata, P. erecta, P. filum, P. flexuosa, P. lescuriana, P. longibracteata, P. longicolla, P. ludwigii, P. melanodon, P. nutans, P. obtusifolia, P. pacifica, P. proligera, P. rabunbaldensis, P. robertsonii, P. tundrae, P. vexans, P. wahlenbergii
P. andalusica, P. andrewsii, P. annotina, P. atropurpurea, P. beringiensis, P. bolanderi, P. brevinervis, P. bulbifera, P. camptotrachela, P. cardotii, P. columbica, P. cruda, P. drummondii, P. elongata, P. erecta, P. filum, P. flexuosa, P. lescuriana, P. longibracteata, P. longicolla, P. ludwigii, P. melanodon, P. nutans, P. obtusifolia, P. pacifica, P. proligera, P. rabunbaldensis, P. robertsonii, P. tundrae, P. vexans, P. wahlenbergii
Synonyms Mnium crudum Bryum crudoides
Name authority (Hedwig) Lindberg: Musc. Scand., 18. (1879) (Sullivant & Lesquereux) Brotherus: in H. G. A. Engler and K. Prantl, Nat. Pflanzenfam. 218[I,3]: 548. (1903)
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