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opal pohlia, opal thread-moss, pohlia moss

pohlia moss

Habit Plants medium-sized to large, green, whitish, or sometimes bluish tinged, shiny. Plants very small to small, whitish, somewhat glossy.
Stems

0.5–2.5 cm.

0.3–1.5 cm.

Leaves

erect to ± spreading, lanceolate to elliptic, 0.8–2 mm;

margins serrulate to serrate in distal 1/3;

costa subpercurrent;

distal medial laminal cells linear-rhomboidal, vermicular, firm, 70–140 µm, walls thin.

erect-spreading, lanceolate, 0.6–1.4 mm;

base not or scarcely decurrent;

margins serrulate in distal 1/3;

costa subpercurrent;

distal medial laminal cells linear-rhomboidal, 60–110 × 6–9 µm, walls thin.

Seta

orange to orange-brown.

orange-brown.

Sexual condition

paroicous, rarely dioicous;

perigonial leaves in dioicous plants linear- or long-lanceolate from ovate base;

perichaetial leaves strongly or rarely weakly differentiated, linear-lanceolate.

dioicous;

perigonial leaves to 3 mm;

perichaetial leaves weakly differentiated, lanceolate.

Capsule

inclined 10–135°, stramineous to orange-brown, long- and slender-pyriform, neck 1/2 urn length;

exothecial cells elongate-rectangular, walls straight;

stomata superficial;

annulus present;

operculum conic;

exostome teeth yellow-brown to red-brown, acute-triangular;

endostome hyaline, basal membrane 1/2 exostome length, segments broadly keeled, broadly perforate, cilia short to long.

inclined ± 180°, brown to stramineous, short-pyriform to urceolate, neck less than 1/3 urn length;

exothecial cells isodiametric, walls sinuate;

stomata superficial or slightly immersed;

annulus absent;

operculum short- to long-conic;

exostome teeth dark to light brown or red-brown, triangular-acute;

endostome light yellow, basal membrane 1/2 exostome length or slightly longer, segments tapered apically, distinctly keeled, broadly perforate, cilia long, nodulose.

Spores

18–26 µm, ± coarsely papillose.

16–21 µm, finely roughened.

Specialized

asexual reproduction absent.

asexual reproduction absent.

Pohlia cruda

Pohlia brevinervis

Phenology Capsules mature summer (Jun–Aug). Capsules mature spring (Apr–Jun).
Habitat Soil banks, rock crevices, under roots, tundra soil and paths Clay soil along streams, ditches, glacial outwash
Elevation moderate to high elevations low to high elevations
Distribution
from FNA
AK; AZ; CA; CO; IA; ID; MA; MD; ME; MI; MN; MT; NC; NE; NJ; NM; NV; NY; OR; PA; SD; TN; UT; VA; VT; WA; WV; WY; AB; BC; MB; NB; NL; NS; NT; NU; ON; QC; SK; YT; Mexico; South America; Australia; Greenland; Pacific Islands; Eurasia; Antarctica
[WildflowerSearch map]
from FNA
AK; AB; YT; Asia (e Siberia)
Discussion

Pohlia cruda is distinguished by the glossy, pale green to whitish or bluish leaves that are narrowly elliptic to lanceolate; the laminal cells are long and narrowly linear-vermicular; the perichaetial leaves are strongly differentiated and linear-lanceolate; the sporophytes have necks about as long as the urn, with long-tapered exostome teeth, broadly keeled and perforate segments, and short to long cilia. Along with P. nutans, P. cruda is one of the most common North American species of the genus. Unlike P. nutans, P. cruda is restricted to northern and montane sites where it grows in rock crevices and on soil banks. The leaves of this species are sometimes slightly complanate. The gametophytes can be unisexual (dioicous) or bisexual (paroicous).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Pohlia brevinervis is a rare and poorly known species that seems to combine characters of P. vexans (erect, glossy leaves on sterile plants) and P. atropurpurea (dark, reddish brown exostome teeth and somewhat immersed stomata). The leaves are distinctly more whitish than those of P. vexans. The exothecial cell walls of capsules of this species are somewhat collenchymatous.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 28, p. 198. FNA vol. 28, p. 209.
Parent taxa Mielichhoferiaceae > Pohlia Mielichhoferiaceae > Pohlia
Sibling taxa
P. andalusica, P. andrewsii, P. annotina, P. atropurpurea, P. beringiensis, P. bolanderi, P. brevinervis, P. bulbifera, P. camptotrachela, P. cardotii, P. columbica, P. crudoides, P. drummondii, P. elongata, P. erecta, P. filum, P. flexuosa, P. lescuriana, P. longibracteata, P. longicolla, P. ludwigii, P. melanodon, P. nutans, P. obtusifolia, P. pacifica, P. proligera, P. rabunbaldensis, P. robertsonii, P. tundrae, P. vexans, P. wahlenbergii
P. andalusica, P. andrewsii, P. annotina, P. atropurpurea, P. beringiensis, P. bolanderi, P. bulbifera, P. camptotrachela, P. cardotii, P. columbica, P. cruda, P. crudoides, P. drummondii, P. elongata, P. erecta, P. filum, P. flexuosa, P. lescuriana, P. longibracteata, P. longicolla, P. ludwigii, P. melanodon, P. nutans, P. obtusifolia, P. pacifica, P. proligera, P. rabunbaldensis, P. robertsonii, P. tundrae, P. vexans, P. wahlenbergii
Synonyms Mnium crudum
Name authority (Hedwig) Lindberg: Musc. Scand., 18. (1879) Lindberg & Arnell: Kongl. Svenska Vetensk. Akad. Handl., n. s. 23(10): 51. (1890)
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