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opal pohlia, opal thread-moss, pohlia moss

Bolander's pohlia moss

Habit Plants medium-sized to large, green, whitish, or sometimes bluish tinged, shiny. Plants small to medium-sized, green or sometimes pale whitish, somewhat shiny.
Stems

0.5–2.5 cm.

0.3–1.5 cm.

Leaves

erect to ± spreading, lanceolate to elliptic, 0.8–2 mm;

margins serrulate to serrate in distal 1/3;

costa subpercurrent;

distal medial laminal cells linear-rhomboidal, vermicular, firm, 70–140 µm, walls thin.

erect to ± spreading, lanceolate to narrowly ovate-lanceolate, 0.8–1.6 mm;

base not or scarcely decurrent;

margins serrulate to serrate in distal 1/3;

costa subpercurrent or percurrent;

distal medial laminal cells linear-rhomboidal, 55–100 µm, walls thin.

Seta

orange to orange-brown.

stramineous to orange-brown.

Sexual condition

paroicous, rarely dioicous;

perigonial leaves in dioicous plants linear- or long-lanceolate from ovate base;

perichaetial leaves strongly or rarely weakly differentiated, linear-lanceolate.

dioicous;

perigonial leaves ovate;

perichaetial leaves slightly differentiated, ovate-lanceolate.

Capsule

inclined 10–135°, stramineous to orange-brown, long- and slender-pyriform, neck 1/2 urn length;

exothecial cells elongate-rectangular, walls straight;

stomata superficial;

annulus present;

operculum conic;

exostome teeth yellow-brown to red-brown, acute-triangular;

endostome hyaline, basal membrane 1/2 exostome length, segments broadly keeled, broadly perforate, cilia short to long.

inclined 35–90°, stramineous to orange-brown, narrowly cylindric, neck 1/2–1 time urn length;

exothecial cells elongate-rectangular, walls straight;

stomata superficial;

annulus present;

operculum conic;

exostome teeth yellow to yellow-brown, acute-triangular;

endostome hyaline, basal membrane 1/4–1/2 exostome length, segments tapered apically, narrowly keeled, narrowly to not perforate, cilia absent to rudimentary.

Spores

18–26 µm, ± coarsely papillose.

16–26 µm, distinctly roughened.

Specialized

asexual reproduction absent.

asexual reproduction absent.

Pohlia cruda

Pohlia bolanderi

Phenology Capsules mature summer (Jun–Aug). Capsules mature summer (Jun–Aug).
Habitat Soil banks, rock crevices, under roots, tundra soil and paths Dry alpine soil, soil-filled rock crevices
Elevation moderate to high elevations low to high elevations
Distribution
from FNA
AK; AZ; CA; CO; IA; ID; MA; MD; ME; MI; MN; MT; NC; NE; NJ; NM; NV; NY; OR; PA; SD; TN; UT; VA; VT; WA; WV; WY; AB; BC; MB; NB; NL; NS; NT; NU; ON; QC; SK; YT; Mexico; South America; Australia; Greenland; Pacific Islands; Eurasia; Antarctica
[WildflowerSearch map]
from FNA
AK; CA; CO; NM; NV; OR; UT; WA; WY; BC; Europe
Discussion

Pohlia cruda is distinguished by the glossy, pale green to whitish or bluish leaves that are narrowly elliptic to lanceolate; the laminal cells are long and narrowly linear-vermicular; the perichaetial leaves are strongly differentiated and linear-lanceolate; the sporophytes have necks about as long as the urn, with long-tapered exostome teeth, broadly keeled and perforate segments, and short to long cilia. Along with P. nutans, P. cruda is one of the most common North American species of the genus. Unlike P. nutans, P. cruda is restricted to northern and montane sites where it grows in rock crevices and on soil banks. The leaves of this species are sometimes slightly complanate. The gametophytes can be unisexual (dioicous) or bisexual (paroicous).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Pohlia bolanderi has glossy to whitish leaves. Glossy forms might be confused with P. longicolla, but P. bolanderi can be distinguished by its smaller size and dioicous inflorescences. The leaves of P. bolanderi are smaller and relatively broader than those of P. longicolla. Variety seriata is characterized by whitish, ranked leaves; it is not recognized here because of intermediate plants from western North America, although in their best development these plants are very distinctive. Plants of this variant might be confused with Conostomum tetragonum (Bartramiaceae).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 28, p. 198. FNA vol. 28, p. 198.
Parent taxa Mielichhoferiaceae > Pohlia Mielichhoferiaceae > Pohlia
Sibling taxa
P. andalusica, P. andrewsii, P. annotina, P. atropurpurea, P. beringiensis, P. bolanderi, P. brevinervis, P. bulbifera, P. camptotrachela, P. cardotii, P. columbica, P. crudoides, P. drummondii, P. elongata, P. erecta, P. filum, P. flexuosa, P. lescuriana, P. longibracteata, P. longicolla, P. ludwigii, P. melanodon, P. nutans, P. obtusifolia, P. pacifica, P. proligera, P. rabunbaldensis, P. robertsonii, P. tundrae, P. vexans, P. wahlenbergii
P. andalusica, P. andrewsii, P. annotina, P. atropurpurea, P. beringiensis, P. brevinervis, P. bulbifera, P. camptotrachela, P. cardotii, P. columbica, P. cruda, P. crudoides, P. drummondii, P. elongata, P. erecta, P. filum, P. flexuosa, P. lescuriana, P. longibracteata, P. longicolla, P. ludwigii, P. melanodon, P. nutans, P. obtusifolia, P. pacifica, P. proligera, P. rabunbaldensis, P. robertsonii, P. tundrae, P. vexans, P. wahlenbergii
Synonyms Mnium crudum Bryum bolanderi, P. bolanderi var. seriata
Name authority (Hedwig) Lindberg: Musc. Scand., 18. (1879) (Lesquereux) Brotherus: in H. G. A. Engler and K. Prantl, Nat. Pflanzenfam. 218[I,3]: 548. (1903)
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