opal pohlia, opal thread-moss, pohlia moss
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pohlia moss
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Plants medium-sized to large, green, whitish, or sometimes bluish tinged, shiny. |
Plants very small to large, as scattered plants or deep, sometimes extensive, turfs. |
0.5–2.5 cm. |
0.1–10 cm, not complanate-foliate, often bearing subfloral innovations below the gametangia. |
erect to ± spreading, lanceolate to elliptic, 0.8–2 mm; margins serrulate to serrate in distal 1/3; costa subpercurrent; distal medial laminal cells linear-rhomboidal, vermicular, firm, 70–140 µm, walls thin. |
with dorsal and lateral not differentiated, erect to spreading, narrowly lanceolate to ovate-lanceolate [rarely linear-lanceolate], 0.6–3.5 mm; base decurrent or not; margins plane (strongly revolute in P. cardotii and P. crudoides), finely serrate to serrulate near apex, rarely entire; apex acute (sometimes short-acuminate in P. melanodon); costa ending well before apex, subpercurrent, percurrent, or rarely short-excurrent; distal medial laminal cells elongate-hexagonal, rhombic, or linear-rhomboidal, sometimes narrowly vermicular; marginal cells undifferentiated or slightly narrower and longer than medial cells. |
orange to orange-brown. |
single (multiple in P. robertsonii), stramineous to orange-brown, straight or bent. |
paroicous, rarely dioicous; perigonial leaves in dioicous plants linear- or long-lanceolate from ovate base; perichaetial leaves strongly or rarely weakly differentiated, linear-lanceolate. |
dioicous, paroicous, or rarely autoicous; perigonia and perichaetia terminal (perigonia rarely lateral in P. longibracteata); perigonial leaf apex acute to long-acuminate; perichaetial leaves same size as vegetative leaves or sometimes strongly differentiated, lanceolate to narrowly linear-lanceolate. |
inclined 10–135°, stramineous to orange-brown, long- and slender-pyriform, neck 1/2 urn length; exothecial cells elongate-rectangular, walls straight; stomata superficial; annulus present; operculum conic; exostome teeth yellow-brown to red-brown, acute-triangular; endostome hyaline, basal membrane 1/2 exostome length, segments broadly keeled, broadly perforate, cilia short to long. |
inclined 0–180°, narrowly cylindric to broadly pyriform or urceolate, 1–6[–10] mm, neck sometimes 1+ times urn length; exothecial cells near mouth in 1–3+ rows, medial cell walls straight or sinuate; stomata superficial to immersed; annulus absent or present, revoluble; operculum convex, short- to long-conic, sometimes short-rostrate; peristome double [rarely absent], 4:2:4–6(–8); exostome white, pale yellow to brown, or sometimes dark reddish brown, teeth triangular to lanceolate, sometimes irregular [rarely reduced or absent], trabeculate or not, pitted basally, coarsely papillose distally (coarsely papillose throughout in P. crudoides); endostome basal membrane low to high, sometimes scarcely exceeding capsule rim, segments narrow to wide, strongly to weakly or scarcely keeled, rarely reduced to rudimentary projections, broadly to narrowly perforate or less often not, cilia absent or 1–3. |
18–26 µm, ± coarsely papillose. |
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asexual reproduction absent. |
asexual reproduction common, by spheric to ovoid rhizoidal tubers or axillary filiform to bulbiform gemmae. |
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Capsules mature summer (Jun–Aug). |
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Soil banks, rock crevices, under roots, tundra soil and paths |
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moderate to high elevations |
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AK; AZ; CA; CO; IA; ID; MA; MD; ME; MI; MN; MT; NC; NE; NJ; NM; NV; NY; OR; PA; SD; TN; UT; VA; VT; WA; WV; WY; AB; BC; MB; NB; NL; NS; NT; NU; ON; QC; SK; YT; Mexico; South America; Australia; Greenland; Pacific Islands; Eurasia; Antarctica
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North America; Mexico; Central America; South America; Europe; Asia; Africa; Pacific Islands (New Zealand); Australia |
Pohlia cruda is distinguished by the glossy, pale green to whitish or bluish leaves that are narrowly elliptic to lanceolate; the laminal cells are long and narrowly linear-vermicular; the perichaetial leaves are strongly differentiated and linear-lanceolate; the sporophytes have necks about as long as the urn, with long-tapered exostome teeth, broadly keeled and perforate segments, and short to long cilia. Along with P. nutans, P. cruda is one of the most common North American species of the genus. Unlike P. nutans, P. cruda is restricted to northern and montane sites where it grows in rock crevices and on soil banks. The leaves of this species are sometimes slightly complanate. The gametophytes can be unisexual (dioicous) or bisexual (paroicous). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Species ca. 85 (32 species in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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1. Axillary gemmae present | → 2 |
1. Axillary gemmae absent | → 14 |
2. Gemmae with peglike leaf primordia | → 3 |
2. Gemmae with broadly laminate (leaflike) leaf primordia | → 8 |
| → 4 |
| → 5 |
4. Gemmae oblong-linear to linear-vermicular, leaf primordia elongate, erect. | P. proligera |
4. Gemmae isodiametric to oblong, leaf primordia inconspicuous, incurved. | P. andrewsii |
5. Gemmae oblong, knobby in outline, leaf primordia absent. | P. flexuosa |
5. Gemmae spheroid, obconic, or linear, not knobby in outline, leaf primordia present | → 6 |
| P. camptotrachela |
6. Gemmae oblong or obconic to linear | → 7 |
7. Gemmae oblong or obconic, leaf primordia 2-5, conspicuous. | P. annotina |
7. Gemmae narrowly linear, leaf primordia 1 or 2, inconspicuous. | P. flexuosa |
8. Gemmae 2-8 per leaf axil, narrowly elongate, oblong-bulbiform, or obconic | → 9 |
8. Gemmae 1 or 2 per leaf axil, bulbiform | → 11 |
9. Gemmae elongate, extending beyond leaves. | P. tundrae |
9. Gemmae oblong, obconic, or spheroid, shorter than leaves | → 10 |
10. Gemmae obconic to oblong. | P. andalusica |
10. Gemmae spheroid to obovate. | P. bulbifera |
11. Gemmae oblong or elliptic to subglobose, leaf primordia small, stiff, restricted to apex or rarely 1 or 2 proximally. | P. filum |
11. Gemmae bulbiform, leaf primordia large, flexuose, at gemma apex and proximally | → 12 |
12. Plants dull; leaves slightly contorted when dry. | P. rabunbaldensis |
12. Plants glossy; leaves unaltered when dry | → 13 |
13. Plants dark green to reddish; leaves ± spreading. | P. drummondii |
13. Plants whitish; leaves ± erect. | P. beringiensis |
14. Leaf margins strongly recurved to revolute | → 15 |
| → 16 |
15. Leaves narrowly long-lanceolate; medial laminal cells linear-hexagonal, walls thin, not porose. | P. crudoides |
15. Leaves ovate-lanceolate; medial laminal cells rhombic, walls thick, ± porose. | P. cardotii |
16. Sexual condition paroicous | → 17 |
16. Sexual condition dioicous (very rarely paroicous) | → 22 |
| P. robertsonii |
| → 18 |
18. Medial laminal cells lax, hexagonal or broadly rhombic, walls thin. | P. obtusifolia |
18. Medial laminal cells firm, hexagonal to rhomboidal or vermicular, walls thin or thick | → 19 |
19. Plants dull; laminal cells hexagonal to rhomboidal | → 20 |
19. Plants shiny; laminal cells linear-vermicular | → 21 |
20. Capsule necks 1/2 urn length; endostome segments broadly perforate, cilia present. | P. nutans |
20. Capsule necks 1/2 to 1 time urn length; endostome segments narrowly to not perforate, cilia absent to rudimentary, rarely long. | P. elongata |
21. Leaves long-lanceolate; endostome segments narrowly to not perforate, cilia absent to rudimentary. | P. longicolla |
21. Leaves lanceolate to elliptic; endostome segments broadly perforate, cilia present, short to long. | P. cruda |
22. Capsules inclined 0-20°; endostome segments weakly keeled, slightly tapered, cilia absent. | P. erecta |
22. Capsules inclined 35-180°; endostome segments distinctly keeled, tapered apically, cilia present or absent | → 23 |
23. Leaf bases long-decurrent; medial laminal cells rhombic to rhomboidal. | P. ludwigii |
23. Leaf bases not or scarcely decurrent; medial laminal cells linear, hexagonal, or rhomboidal | → 24 |
24. Capsules narrowly cylindric, inclined 35-90°; endostome segments narrowly to not perforate, cilia absent to rudimentary. | P. bolanderi |
24. Capsules pyriform to urceolate, inclined 80-180°; endostome segments broadly perforate, cilia present | → 25 |
25. Annulus present; exothecial cells rectangular | → 26 |
25. Annulus absent; exothecial cells isodiametric | → 28 |
26. Plants green or rarely reddish, dull; laminal cells short- hexagonal. | P. nutans |
26. Plants pale green to green, glossy; laminal cells linear-rhomboidal to linear-vermicular | → 27 |
27. Plants somewhat glossy; leaf margins serrulate at apex; medial laminal cells 65-110 µm. | P. lescuriana |
27. Plants very glossy; leaf margins coarsely serrate at apex; medial laminal cells 110-145 µm. | P. pacifica |
28. Medial laminal cells laxly rhomboidal | → 29 |
28. Medial laminal cells linear-hexagonal to linear-rhomboidal or broadly rhomboidal | → 31 |
29. Plants very small, reddish; stems 0.2-0.4 cm; stomata superficial. | P. atropurpurea |
29. Plants small to large, green to whitish, rarely reddish when very large; stems 0.3-10 cm; stomata deeply immersed | → 30 |
30. Plants dark green; stems 0.3-1 cm. | P. melanodon |
30. Plants whitish, sometimes reddish when very large; stems 0.8-10 cm. | P. wahlenbergii |
31. Plants medium-sized to large; leaves wide-spreading; perigonial leaves to 6 mm. | P. longibracteata |
31. Plants small to rarely medium-sized; leaves erect to erect-spreading; perigonial leaves to 3 mm | → 32 |
32. Stomata superficial or slightly immersed | → 33 |
| → 34 |
33. Stems 0.3-1.5 cm; leaves erect-spreading; medial laminal cells linear, 6-9 µm wide. | P. brevinervis |
33. Stems 0.2-0.4 cm; leaves erect-spreading to spreading; medial laminal cells laxly and broadly rhomboidal, 10-12 µm wide. | P. atropurpurea |
34. Stems green to pale orange or pink; leaves spreading; plants dull. | P. columbica |
34. Stems cherry red; leaves erect; plants glossy. | P. vexans |
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FNA vol. 28, p. 198. |
FNA vol. 28, p. 193. |
Mielichhoferiaceae > Pohlia |
Mielichhoferiaceae |
P. andalusica, P. andrewsii, P. annotina, P. atropurpurea, P. beringiensis, P. bolanderi, P. brevinervis, P. bulbifera, P. camptotrachela, P. cardotii, P. columbica, P. crudoides, P. drummondii, P. elongata, P. erecta, P. filum, P. flexuosa, P. lescuriana, P. longibracteata, P. longicolla, P. ludwigii, P. melanodon, P. nutans, P. obtusifolia, P. pacifica, P. proligera, P. rabunbaldensis, P. robertsonii, P. tundrae, P. vexans, P. wahlenbergii |
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P. andalusica, P. andrewsii, P. annotina, P. atropurpurea, P. beringiensis, P. bolanderi, P. brevinervis, P. bulbifera, P. camptotrachela, P. cardotii, P. columbica, P. cruda, P. crudoides, P. drummondii, P. elongata, P. erecta, P. filum, P. flexuosa, P. lescuriana, P. longibracteata, P. longicolla, P. ludwigii, P. melanodon, P. nutans, P. obtusifolia, P. pacifica, P. proligera, P. rabunbaldensis, P. robertsonii, P. tundrae, P. vexans, P. wahlenbergii |
Mnium crudum |
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(Hedwig) Lindberg: Musc. Scand., 18. (1879) |
Hedwig: Sp. Musc. Frond., 171. (1801) |
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