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panic grass, perennial panicgrass, rosette grass

Habit Plants annual or perennial; habit various. Plants perennial; cespitose, sometimes rhizomatous, sometimes with hard, cormlike bases, often with basal winter rosettes of leaves having shortly ovate to lanceolate blades, these often sharply distinct from the blades of the cauline leaves.
Culms

3-800 cm, annual, usually not woody.

5-150 cm, herbaceous, hollow, usually erect or ascending, rarely sprawling, in the spring often spreading, sometimes decumbent in the fall, usually branching from the mid- or lower culm nodes in summer and fall;

branches rebranching 1-4 times, terminating in small secondary panicles that are usually partly included in the sheaths.

Leaves

basal and/or cauline;

sheaths usually open;

ligules of hairs or membranous, membranous ligules often ciliate, cilia sometimes longer than the membranous base;

blades occasionally pseudopetiolate, seldom disarticulating at maturity.

Cauline leaves

3-14, usually distinctly longer and narrower than the rosette blades;

ligules of hairs, membranous, or membranous and ciliate, sometimes absent;

pseudoligules of 1-5 mm hairs often present at the bases of the blades immediately behind the true ligules;

blades usually distinctly longer and narrower than those of the basal rosette, cross sections with non-Kranz anatomy;

photosynthesis C3.

Inflorescences

terminal, sometimes also axillary, occasionally subterranean panicles;

branches sometimes spikelike and secund, sometimes less than 1 cm;

disarticulation usually below the glumes, sometimes at the base of the panicle branches, occasionally below the florets.

panicles, terminal on the culms and branches;

sterile branches and bristles absent;

disarticulation below the glumes.

Spikelets

usually dorsally compressed, varying to terete or laterally compressed, with 2(3) florets, lower florets staminate, sterile, or reduced, upper florets usually bisexual;

calluses not developed.

0.8-5.2 mm, not subtended by bristles, dorsally compressed, surfaces unequally convex, apices unawned.

Glume(s)

usually membranous;

lower glumes usually less than 1/2 as long as the spikelets, sometimes absent;

upper glumes usually subequal to the upper florets, occasionally absent;

lower lemmas similar to the upper glumes in length and texture;

upper lemmas indurate, coriaceous, or cartilaginous, with a germination flap at the base, margins usually widely separated and involute at maturity, sometimes flat and hyaline;

upper paleas similar to the upper lemmas in length and texture;

lodicules short;

anthers usually 3;

stigmas usually red.

apices not or only slightly gaping at maturity;

lower glumes 1/5 – 3/4 as long as the spikelets, 1-5-veined, truncate, acute, or acuminate;

upper glumes slightly shorter than the spikelets or exceeding the upper florets by up to 1 mm, 5-11-veined, not saccate, apices rounded to attenuate.

Caryopses

usually dorsally compressed or terete;

embryos 1/2 or more the length of the caryopses.

smooth;

pericarp thin;

endosperm hard;

hila round or oval, x = 9.

Primary

panicles terminating the culms, developing April-June (July), sometimes also in late fall, usually at least partially chasmogamous, often with a lower seed set than the secondary panicles;

secondary panicles terminating the branches, produced from (May)June to fall, usually partially or totally cleistogamous.

Lower

florets sterile or stami-nate;

lower lemmas similar to the upper glumes;

lower paleas sometimes present, thin, shorter than the lower lemmas;

upper florets bisexual, sessile, plump, usually apiculate to mucronate, sometimes minutely so, or subacute to (rarely) acute;

upper lemmas striate, chartaceous-indurate, shiny, usually glabrous, margins involute;

upper paleas striate;

lodicules 2;

anthers 3.

x

= 9, 10.

Poaceae tribe Paniceae

Dichanthelium

Distribution
from FNA
AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; HI; PR; AB; BC; LB; MB; NB; NS; ON; PE; QC; SK
[BONAP county map]
Discussion

The tribe Paniceae, which includes about 100 genera and 2000 species, is primarily tropical in distribution. Within the Flora region, it is represented by 27 genera and 262 species, with its greatest representation being in the eastern portion of the contiguous United States (Barkworth and Capels 2000).

The tribe is so morphologically distinct that it was first recognized, in essentially its current sense, by Robert Brown in 1814. Its primary distinguishing features are the unusual spikelet structure combined with the indurate to coriaceous upper florets. Recent molecular studies (Barber et al. 2002; Guissani et al. 2001) show it as comprising two distinct lineages, one of which contains species with a base number of x = 9 and the other, species with x = 10.

Photosynthesis in the Paniceae may follow the C3 pathway or any of three different C4 pathways. Most genera are uniform in this regard, but there are some noteworthy exceptions. Guissani et al. (2001) concluded that the C3 pathway is probably ancestral within the tribe and that two of the three C4 pathways, NAD-ME and PCK, originated only once within the tribe, whereas the NADP-ME pathway originated independently in several different lineages. Most genera are uniform with respect to their photosyntheticd pathway, but there are some noteworthy exceptions.

The germination flap is a small area of soft tissue at the base of the upper lemma through which the primary root of the seedling grows.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Dichanthelium is a genus of approximately 72 species, 34 of which are native to the Flora region. It is often included in Fanicum, the two taxa being similar in gross morphology. Recent molecular data reinforce the morphological arguments for recognizing Dichanthelium as a distinct genus.

When the branches of Dichanthelium develop, in late summer or fall, the culms acquire a very different aspect; comments about the 'fall phase' refer to the appearance of the plant or its culms following this branching. Unless stated otherwise, descriptions and measurements refer to structures of the culms and primary panicles, not those of the branches and secondary panicles. Ligule measurements usually include the hairs of the pseudoligule, if present, because the two are often difficult to distinguish with less than 30x magnification.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Plants developing both subterranean and aerial inflorescences, only the subterranean spikelets setting seed
Amphicarpum
1. Plants developing only aboveground inflorescences, the spikelets setting seed [Amphicarpum is also keyed out here to accommodate situations in which looking for subterranean inflorescences is not permitted or specimens have no underground parts].
→ 2
2. Inflorescences spikelike panicles, with the branches partially embedded in the flattened rachises; plants perennial, stoloniferous
Stenotaphrum
2. Inflorescences panicles, sometimes spikelike, but the branches not embedded in the rachises or the rachises not flattened; plants annual or perennial, sometimes stoloniferous.
→ 3
3. Most spikelets or groups of 2-11 spikelets subtended by 1-many, distinct to more or less connate, stiff bristles or bracts.
→ 4
4. Spikelets in groups of 2-11, subtended by 4 flat, narrowly elliptic, coriaceous bracts; terete bristles not present
Anthephora
4. Spikelets solitary or in groups, subtended by 1-many stiff, terete bristles, sometimes appearing as an extension of the branch; flat, connate bristles sometimes present distal to the terete bristles.
→ 5
5. Bristles falling with the spikelets at maturity; disarticulation at the base of the reduced panicle branches (fascicles).
→ 6
6. Bristles plumose or antrorsely scabrous, free or fused no more than 1/2 their length
Pennisetum
6. Bristles glabrous, smooth, retrorsely scabrous, or strigose, usually at least some bristles fused for more than 1/2 their length
Cenchrus
5. Bristles persistent; disarticulation below the spikelets.
→ 7
7. Upper glumes indurate at maturity; lower lemmas somewhat indurate at the base; pedicels subtended by a single bristle
Setariopsis
7. Upper glumes membranous to herbaceous at maturity; lower lemmas neither constricted nor indurate at the base; pedicels subtended by 1-many bristles.
→ 8
8. Spikelets subtended by 1-many bristles; paleas of the lower florets usually hyaline to membranous at maturity, rarely absent or reduced; paleal veins not keeled
Setaria
8. Spikelets subtended by 1 bristle; paleas of the lower florets coriaceous to indurate at maturity, the keels thickened
Ixophorus
3. All or most spikelets not subtended by stiff bristles, sometimes the terminal spikelet on each branch subtended by a single bristle, and occasionally other spikelets with a single subtending bristle.
→ 9
9. Terminal spikelet on each branch subtended by a single bristle; other spikelets occasionally with a single stiff subtending bristle
Setaria
9. None of the spikelets subtended by a stiff bristle.
→ 10
10. Inflorescences of spikelike branches 1-3.7 cm long, the branch axes extending as a 2.5-4 mm bristle beyond the base of the distal spikelets
Paspalidium
10. Inflorescences various but, if of spikelike branches, these terminating in a well-developed or rudimentary spikelet.
→ 11
11. Lower glumes or lower lemmas awned, sometimes shortly so (the awn reduced to a point in Echinochloa colona).
→ 12
12. Upper florets laterally compressed; spikelets also laterally compressed
Melinis
12. Upper florets dorsally compressed; spikelets usually dorsally compressed or terete, sometimes laterally compressed.
→ 13
13. Blades linear to linear-lanceolate, usually more than 10 times longer than wide, with prominent midribs; at least the upper leaves, often all leaves, without ligules; ligules usually absent, particularly from the upper leaves, of hairs when present
Echinochloa
13. Blades triangular to lanceolate, less than 10 times longer than wide, the midribs not particularly prominent, at least distally; ligules present, of hairs or membranous.
→ 14
14. Lower glumes awned, the awns exceeding the florets; upper glumes not ciliate-margined; culms trailing on the ground, frequently rooting and branching at the nodes
Oplismenus
14. Lower glumes unawned or shortly awned, the awns exceeded by the florets; upper glumes ciliate-margined; culms erect or decumbent below, sometimes rooting and branching at the lower nodes
Alloteropsis
11. Lower glumes and lower lemmas unawned.
→ 15
15. Upper florets laterally compressed
Melinis
15. Upper florets dorsally compressed or terete.
→ 16
16. Upper lemmas and paleas cartilaginous and flexible at maturity; lemma margins flat, hyaline; lower glumes absent or to 1/4 the length of the spikelets.
→ 17
17. Aerial inflorescences with elongate rachises and glabrous spikelets; spikelets of the aerial panicles rarely setting seed; subterranean spikelets developed, seed-forming
Amphicarpum
17. Aerial inflorescences of digitate or subdigitate clusters of spikelike branches with glabrous or pubescent spikelets or with elongate rachises and conspicuously pubescent spikelets; aerial spikelets seed-forming; subterranean spikelets not developed.
→ 18
18. Spikelets ellipsoid to obovoid; inflorescences simple panicles with erect to ascending branches on elongate rachises; branches ascending, not conspicuously spikelike
Anthenantia
18. Spikelets lanceoloid to ellipsoid; inflorescences usually panicles with digitate or subdigitate clusters of spikelike branches, sometimes simple panicles with strongly divergent branches
Digitaria
16. Upper lemmas and paleas chartaceous to indurate and rigid at maturity; lemma margins not hyaline, frequently involute; lower glumes varying from absent to subequal to the spikelets or extending beyond the distal floret.
→ 19
19. Spikelets subtended by a cuplike callus
Eriochloa
19. Spikelets not subtended by a cuplike callus.
→ 20
20. At least the upper leaves, often all leaves, without ligules; ligules, when present, of hairs
Echinochloa
20. All leaves with ligules, ligules membranous or of hairs.
→ 21
21. Paleas of the lower florets inflated and indurate at maturity; lower and upper florets standing apart from each other when mature
Steinchisma
21. Paleas of the lower florets neither inflated nor indurate at maturity; lower and upper florets closely appressed to each other when mature, [revert to left side]
→ 22
22. Inflorescences of 1-sided, spikelike primary branches.
→ 23
23. Spikelets with the lower lemmas and lower glumes (if present) adjacent to the branch axes.
→ 24
24. Lower glumes absent
Axonopus
24. Lower glumes present on all or most spikelets.
→ 25
25. Upper lemmas rugose and verrucose; panicle branches in 2 or more ranks, sometimes verticillate
Urochloa
25. Upper lemmas smooth; panicle branches in 1 rank
Moorochloa
23. Spikelets with the upper lemmas and upper glumes (if present) adjacent or appressed to the branch axes.
→ 26
26. Both glumes absent from all or almost all spikelets, the terminal spikelet on a branch sometimes with upper glumes
Reimarochloa
26. Upper or both glumes present on all spikelets.
→ 27
27. Upper lemmas smooth to slightly rugose; lower glumes usually absent
Paspalum
27. Upper lemmas rugose and verrucose; lower glumes present, from 1/5 as long as the spikelets to equaling them
Urochloa
22. Inflorescences usually panicles with well-developed secondary branches, sometimes spikelike panicles or panicles with spikelike, but not 1-sided, branches.
→ 28
28. Inflorescences dense, the spikelets concealing at least the distal 1/2 of the rachises.
→ 29
29. Upper glumes slightly to strongly saccate, 5-13-veined; panicle branches often fused to the rachises; blades 1.5-22 mm wide; culm internodes hollow
Sacciolepis
29. Upper glumes not saccate, 3-7-veined; panicle branches not fused to the rachises; blades 12-28 mm wide; culm internodes filled with aerenchyma
Hymenachne
28. Inflorescences more or less open panicles, the spikelets not concealing the rachises.
→ 30
30. Lower glumes with saccate bases; glumes and lemmas with woolly pubescent apices; culms weakly lignified, rooting at the nodes
Lasiacis
30. Lower glumes not saccate basally; glumes and lemmas glabrous or with short, straight hairs, apices sometimes with a tuft of hairs but never woolly pubescent; culms usually not lignified, if lignified, not rooting at the nodes.
→ 31
31. Lemmas of the upper florets rugose and verrucose; panicle branches usually spikelike and 1-sided, alternate or subopposite, less frequently verticillate
Urochloa
31. Lemmas of the upper florets usually smooth, if rugose the panicle branches neither verticillate nor 1-sided and spikelike.
→ 32
32. Plants developing aerial and subterranean panicles; aerial spikelets lanceoloid, often without lower glumes; upper lemmas with flat margins
Amphicarpum
32. Plants developing aerial, but not subterranean, panicles; spikelets ovoid to ellipsoid or lanceoloid; lower glumes present; upper lemmas with involute margins.
→ 33
33. Blades of the basal leaves clearly distinct from the cauline leaves; basal leaves ovate to lanceolate, cauline leaves with longer and narrower blades; basal leaves forming a distinct winter rosette
Dichanthelium
33. Blades of the basal and cauline leaves similar, usually linear to lanceolate, varying from filiform to ovate; basal leaves not forming a distinct winter rosette.
→ 34
34. Panicles terminating the culms usually appearing in late spring; branches usually developing from the lower and middle cauline nodes in summer, the branches rebranching 1 or more times by fall; upper florets not disarticulating at maturity, plump
Dichanthelium
34. Panicles terminating the culms usually appearing after midsummer; branches usually not developing branches from the lower and middle cauline nodes, when present, rarely rebranched; upper florets disarticulating or not very plump at maturity
Panicum
1. Basal leaf blades similar in shape to those of the lower cauline leaves, usually erect to ascending, clustered at the base, sometimes small or vestigial; culms branching from near the base in the fall, with 2-4 leaves, only the upper 2-4 internodes elongated.
→ 2
2. Blades soft, 3-12 mm wide, usually ciliate; upper blades less than 20 times as long as wide; fall phase with short panicle-bearing branches, without sterile shoots (sect. Strigosa).
→ 3
3. Leaf sheaths with retrorse or spreading hairs; upper blades 4-17 cm long, at least 3/4 as long as the basal blades; blade margins usually finely short ciliate, the cilia not papillose-based; spikelets with papillose-based hairs
D. laxiflorum
3. Leaf sheaths glabrous or with ascending hairs; upper blades 1.5-6 cm long, less than 3/4 as long as the basal blades; blade margins with papillose-based cilia; spikelets glabrous or pubescent, hairs not papillose-based
D. strigosum
2. Blades stiff, 1-5 mm wide, not ciliate; most upper blades at least 20 times as long as wide; fall phase with basal panicles and sterile shoots (sect. Linearifolia).
→ 4
4. Upper glumes and lower lemmas forming a beak extending 0.2-1 mm beyond the upper florets; spikelets 3.2-4.3 mm long; primary panicles with 7-25 spikelets
D. depauperatum
4. Upper glumes and lower lemmas equaling or exceeding the upper florets by no more than 0.3 mm, not forming a beak; spikelets 2-3.4 mm long; primary panicles with 12-70 spikelets.
→ 5
5. Cauline blades 4-8 cm long, all alike; basal blades ascending to spreading
D. wilcoxianum
5. Uppermost cauline blades 10-20 cm long, distinctly longer than the lower blades; basal blades erect to ascending.
→ 6
6. Panicles 1-3 cm wide, with ascending branches and appressed pedicels; spikelets turgid, 2.6-3.4 mm long, 1-1.7 mm wide, upper florets obovoid
D. perlongum
6. Panicles 2-6 cm wide, with spreading branches and pedicels; spikelets not turgid, 2-3.2 mm long, 0 8-1.4 mm wide, upper florets ellipsoid
D. linearifolium
1. Basal leaf blades usually well-differentiated from the cauline blades, ovate to lanceolate, spreading, forming a rosette, or basal blades absent; culms usually branching from the midculm nodes in the fall, with 3-14 leaves, usually all internodes elongated.
→ 7
7. Bases of the culms hard, cormlike; basal rosettes absent; spikelets with papillose-based hairs and attenuate basally (sect. Pedicellata).
→ 8
8. Culms erect in the spring; cauline leaves 4-7, with thin, glabrous or sparsely hirsute blades that widen distal to the rounded to subcordate bases; lower glumes not encircling the pedicels, subadjacent to the upper glumes
D. pedicellatum
8. Culms decumbent to ascending in the spring; cauline leaves 8-14, with thick, firm, puberulent blades that are parallel-sided distal to the rounded to truncate bases; lower glumes almost to completely encircling the pedicels, attached about 0.2 mm below the upper glumes
D. nodatum
7. Bases of the culms not cormlike; basal rosettes usually present; spikelets not both with papillose-based hairs and attenuate basally.
→ 9
9. Blades cordate, thick, with white, cartilaginous margins; spikelets usually spherical to broadly obovoid or broadly ellipsoid, 1-1.8 mm long (sect. Sphaerocarpa).
→ 10
10. Spikelets 1-1.4 mm long; lower glumes 0.2-0.4 mm long; cauline blades 5-10 mm wide
D. erectifolium
10. Spikelets 1.3-1.8 mm long; lower glumes 0.4-0.8 mm long; cauline blades 5-25 mm wide.
→ 11
11. Cauline blades 4-7,10-25 cm long, 14-25 mm wide, with evident veins; culms nearly erect; panicles less than 1/2 as wide as long
D. polyanthes
11. Cauline blades 3-4(6), 1.5-10 cm long, 5-14 mm wide, with obscure veins; culms decumbent or ascending; panicles more than 1/2 as wide as long
D. sphaerocarpon
9. Blades not cordate or the spikelets not both spherical and less than 1.9 mm long; blade margins usually not white and cartilaginous.
→ 12
12. Lower glumes thinner and more weakly veined than the upper glumes, attached about 0.2 mm below the upper glumes, the bases clasping the pedicels; spikelets attenuate basally.
→ 13
13. Blades 2-7 cm long, about 10 times as long as wide, not or slightly involute, spreading, without raised veins, not longitudinally wrinkled; spikelets obovoid-obpyriform, planoconvex in side view (sect. Lancearia)
D. portoricense
13. Blades 4-16 cm long, more than 14 times as long as wide, or involute, stiffly erect or ascending, with prominently raised veins, the lower blades usually longitudinally wrinkled; spikelets ellipsoid to obovoid, biconvex in side view (sect. Angustifolia).
→ 14
14. Culms densely villous; nodes densely bearded; spikelets densely pubescent
D. consanguineum
14. Culms glabrous, puberulent, or pilose with papillose-based hairs; nodes glabrous, puberulent to lightly bearded; spikelets glabrous or pubescent
D. aciculare
12. Lower glumes similar in texture and vein prominence to the upper glumes, attached immediately below the upper glumes, the bases not clasping the pedicels; spikelets usually not attenuate basally.
→ 15
15. Culms arising from rhizomes 3-5 mm thick, with (5)7-14 cauline blades; sheaths strongly hispid or viscid, mottled with pale spots, constricted at the top (sect. Clandestina).
→ 16
16. Nodes densely bearded above a viscid glabrous ring, often swollen; blades densely soft pubescent
D. scoparium
16. Nodes glabrous or sparsely pubescent, not swollen; blades glabrous or sparsely pubescent.
→ 17
17. Cauline blades 7-15 mm wide, apices involute, long tapering; spikelets glabrous or sparsely puberulent
D. scabriusculum
17. Cauline blades 15-30 mm wide, apices flat, acuminate; spikelets sparsely pubescent
D. clandestinum
15. Culms arising from caudices or from rhizomes to 2 mm thick, with 3-7(9) cauline blades; sheaths not viscid, rarely hispid, not mottled with pale spots or constricted at the top.
→ 18
18. Ligules with a membranous base, ciliate distally; culms usually arising from slender rhizomes; lower florets often staminate; cauline blades 5-40 mm wide, often with a cordate base (sect. Macrocarpa).
→ 19
19. Spikelets ellipsoid, not turgid, with pointed apices; cauline blades 4-6, cordate at the base; sheaths without papillose-based hairs.
→ 20
20. Spikelets 2.2-3.2 mm long; ligules about 0.3 mm long; blades 5-25 mm wide; lower floret sterile
D. commutatum
20. Spikelets 2.9-5.2 mm long; ligules 0.4-0.9 mm long; blades 15-40 mm wide; at least some lower florets staminate.
→ 21
21. Nodes glabrous or slightly bearded; spikelets 2.9-3.9 mm long
D. latifolium
21. Nodes densely retrorsely bearded; spikelets 3.8-5.2 mm long
D. boscii
19. Spikelets obovoid, turgid, with rounded apices; cauline blades 3-4, tapered, rounded or truncate to cordate at the base; sheaths with papillose-based hairs.
→ 22
22. Blades and spikelets with papillose-based hairs; panicles usually slightly longer than wide, with spreading to ascending branches
D. leibergii
22. Blades glabrous; spikelets puberulent to almost glabrous; panicles usually more than twice as long as wide, with nearly erect branches
D. ×anthophysum
18. Ligules of hairs (except for D. nudicaule); culms arising from caudices; lower florets sterile; cauline blades 1-18 mm wide, bases usually tapered, rounded, or truncate at the base, sometimes cordate.
→ 23
23. Lower internodes short, upper nodes elongated; flag leaves distant and much reduced; culms rarely branching in the fall; branches, if present, few, developing from basal and subbasal nodes, erect (sect. Nudicaulia)
D. nudicaule
23. Lower internodes about as long as the upper internodes; flag leaves usually not much reduced; culms branching in the fall; branches often many, developing from mid- or upper culm nodes, often spreading.
→ 24
24. Spikelets 2.5-4.3 mm long, usually obovoid, turgid; upper glumes usually with an orange or purple spot at the base, the veins prominent (sect. Oligosantha).
→ 25
25. Nodes glabrous or sparsely pubescent; abaxial surfaces of the blades glabrous or pubescent, but not velvety pubescent
D. oligosanthes
25. Nodes bearded with spreading to retrorse hairs; abaxial surfaces of the blades softly velvety pubescent.
→ 26
26. Spikelets 3.7-4.3 mm long; culms 2-3 mm thick, stiffly erect; ligules 2-5 mm long, without pseudoligules; blades glabrous or sparsely pilose on the adaxial surfaces
D. ravenelii
26. Spikelets 2.5-3.2 mm long; culms usually 1-2 mm thick, erect; ligules 0.5-1 mm long, with the adjacent pseudoligules 1-3 mm long; blades densely velvety pubescent on both surfaces
D. malacophyllum
24. Spikelets 0.8-3 mm long, ellipsoid or obovoid, not turgid; upper glumes lacking an orange or purple spot at the base and the veins not prominent.
→ 27
27. Ligules and adjacent pseudoligules 1-5 mm long, or the culms and sheaths with long hairs and also puberulent; spikelets variously pubescent to subglabrous (sect. Lanuginosa).
→ 28
28. Spikelets 0.8-1.1 mm long, puberulent to subglabrous; culms delicate, 0.3-0.8 mm thick
D. wrightianum
28. Spikelets 1.1-3 mm long, variously pubescent; culms not delicate, usually more than 1 mm thick.
→ 29
29. Spikelets 1.1-2.1 mm long; sheaths glabrous or pubescent with hairs no more than 3 mm long
D. acuminatum
29. Spikelets 1.8-3 mm long; sheaths with hairs to 4 mm long
D. ovale
27. Ligules absent or to 1.8 mm long, without adjacent pseudoligules; culms and at least the upper sheaths glabrous or sparsely pubescent with hairs of 1 length only; spikelets glabrous or pubescent.
→ 30
30. Culms (18)40-100 cm tall, rarely delicate, usually more than 1 mm thick; spikelets 1.5-2.7 mm long; blades 3.5-14 cm long, 5-14 mm wide (sect. Dichanthelium.).
→ 31
31. Spikelets glabrous or, if pubescent, either the nodes bearded or the culms weak and prostrate; blade of the flag leaf usually spreading
D. dichotomum
31. Spikelets pubescent; nodes glabrous; culms erect or ascending; blade of the flag leaf erect or ascending
D. boreale
30. Culms 5-40(55) cm tall, delicate, usually less than 1 mm thick; spikelets 1.1-1.7 mm long; longest blades 1.5-6 cm long, 1.5-6 mm wide (sect. Ensifolia).
→ 32
32. Culms reclining or weakly erect; cauline blades 4-9, usually without prominent white, cartilaginous margins; ligules often more than 1 mm long
D. ensifolium
32. Culms erect, sometimes geniculate basally; cauline blades 3-5, with prominent white, cartilaginous margins; ligules 0.2-0.7 mm long.
→ 33
33. Culms few per clump; the fall phase branching sparingly; cauline blades flat, the bases rounded; blades of the flag leaves much shorter than those of the lower leaves
D. tenue
33. Culms many per clump; the fall phase branching extensively; cauline blades often involute, the bases subcordate; blades of the flag leaves only slightly shorter than those of the lower leaves
D. chamaelonche
Source FNA vol. 25, p. 353. Author: Mary E. Barkworth;. FNA vol. 25, p. 406.
Parent taxa Poaceae > subfam. Panicoideae Poaceae > subfam. Panicoideae > tribe Paniceae
Subordinate taxa
Alloteropsis, Amphicarpum, Anthenantia, Anthephora, Axonopus, Cenchrus, Dichanthelium, Digitaria, Echinochloa, Eriochloa, Hymenachne, Ixophorus, Lasiacis, Melinis, Moorochloa, Oplismenus, Panicum, Paspalidium, Paspalum, Pennisetum, Reimarochloa, Sacciolepis, Setaria, Setariopsis, Steinchisma, Stenotaphrum, Urochloa
D. aciculare, D. acuminatum, D. boreale, D. boscii, D. chamaelonche, D. clandestinum, D. commutatum, D. consanguineum, D. depauperatum, D. dichotomum, D. ensifolium, D. erectifolium, D. latifolium, D. laxiflorum, D. leibergii, D. linearifolium, D. malacophyllum, D. nodatum, D. nudicaule, D. oligosanthes, D. ovale, D. pedicellatum, D. perlongum, D. polyanthes, D. portoricense, D. ravenelii, D. scabriusculum, D. scoparium, D. sphaerocarpon, D. strigosum, D. tenue, D. wilcoxianum, D. wrightianum, D. ×anthophysum
Name authority R. Br. (Hitchc. & Chase) Gould
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