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nodding bluegrass

Hooker's blue grass, veiny bluegrass, Wheeler bluegrass

Habit Plants perennial, short-lived; densely tufted, tuft bases narrow or not, not stoloniferous, not rhizomatous. Plants perennial; loosely tufted or with solitary shoots, short-rhizomatous.
Culms

10-60 cm.

20-65 cm, erect or the bases decumbent, terete or weakly compressed;

nodes terete, 1-2 exserted.

Sheaths

closed for 1/3 – 2/3 their length, terete, smooth;

ligules 1.5-3.5 mm, smooth or sparsely scabrous;

blades 1.5-4 mm wide, flat, thin, soft, apices broadly prow-shaped.

closed for 2/3 – 9/10 their length, terete to slightly compressed, smooth or sparsely scabrous, sometimes hairy, hairs about 0.15 mm, bases of basal sheaths glabrous, distal sheath lengths (0.7)1-2.2(2.8) times blade lengths;

collars of proximal leaves usually hairy on and near the margins, marginal hairs longer than those of the sheaths;

ligules 0.5-1.5 mm, smooth or scabrous, hairy, hairs about 0.15 mm, truncate to obtuse, those of the lower culm and innovation leaves 0.5-1 mm, scabrous or softly puberulent, truncate;

innovation blades similar to or longer than the cauline blades;

cauline blades 2-4.5 mm wide, usually flat, lax, adaxial surfaces smooth or sparsely scabrous, particularly over the veins, apices broadly prow-shaped, blades gradually reduced in length distally or the middle blades longest, flag leaf blades 3-8 cm long.

Basal branching

mixed intra- and extravaginal.

mainly extravaginal.

Panicles

4-15 cm, nodding, open, with numerous spikelets and 1-2 branches per node;

branches (2)3-7 cm, spreading to reflexed, lower branches usually reflexed, flexuous, usually terete, smooth or sparsely scabrous, with (3)6-18 spikelets.

8-15 cm, erect or lax, ovoid to pyramidal, open or loosely contracted, sparse, with 25-80 spikelets, proximal internodes 1.8-3.5 cm;

nodes with 3-5 branches;

branches 2.5-8 cm, ascending to spreading, lax, terete to weakly angled, moderately scabrous, with 2-8 spikelets.

Spikelets

4-6 mm, lanceolate to broadly lanceolate, usually partly to wholly purplish, with 3-5 florets;

rachilla internodes shorter than 1 mm, smooth.

4-7 mm, lengths to 3.5 times widths, laterally compressed, not sexually dimorphic;

florets 3-8;

rachilla internodes smooth or scabrous, glabrous or sparsely hispidulous.

Glumes

narrowly to broadly lanceolate, distinctly keeled, keels smooth or nearly so;

lower glumes 1-veined;

upper glumes shorter than or subequal to the lowest lemmas;

calluses webbed;

lemmas 2-3.5 mm, lanceolate, partly purple to fairly strongly purple, distinctly keeled, keels and marginal veins short- to long-villous, keels hairy for 2h-ls their length, lateral veins usually sparsely softly puberulent at least on 1 side, lateral veins obscure to moderately prominent, intercostal regions smooth, minutely bumpy, glabrous, apices acute, slightly bronze-colored or not;

palea keels scabrous, usually softly puberulent at midlength;

anthers 0.6-1 mm.

2/3 – 4/5 as long as the adjacent lemmas, lanceolate, distinctly keeled;

lower glumes 1-3(5)-veined;

calluses usually glabrous, rarely minutely webbed;

lemmas 3-4.5 mm, lanceolate, distinctly keeled, keels and marginal veins usually glabrous, infrequently sparsely softly puberulent to short-villous, intercostal regions glabrous or hispidulous, smooth or finely scabrous, margins glabrous, apices acute;

paleas scabrous over the keels, intercostal regions glabrous;

anthers usually 2.5-4 mm, sometimes vestigial (0.1-0.2 mm).

2n

= 28.

= 28, 28+1.

Poa reflexa

Poa nervosa

Distribution
from FNA
AZ; CO; ID; MT; NM; NV; OR; UT; WY
[WildflowerSearch map]
[BONAP county map]
from FNA
OR; WA; AB; BC
[WildflowerSearch map]
[BONAP county map]
Discussion

Poa reflexa grows in subalpine forests, meadows, and low alpine habitats, primarily in the central and southern Rocky Mountains. It usually grows on drier and more disturbed sites, and appears shorter-lived, than the frequently sympatric or parapatric P. leptocoma (p. 573), from which it differs in usually having hairs on the palea keels and lateral veins of the lemmas, and smooth panicle branches. In addition, P. reflexa is tetraploid, whereas P. leptocoma is hexaploid. Poa reflexa may resemble small plants of P. occidentalis (see previous) in habit.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Poa nervosa occurs infrequently at low elevations in the western foothills of the northern Cascade Mountains and adjacent coast ranges, extending eastward up the Columbia Gorge as far as Multnomah Falls. It usually grows in wet habitats, such as mossy cliffs with seeps and around waterfalls, but it is also found in rich, old growth, mixed deciduous and conifer forests. It appears to be sexually reproducing and sequentially gynomonoecious.

Poa nervosa differs from P. wheeleri (see next) in having densely pubescent leaf collar margins, and glabrous or more sparsely and shortly pubescent sheaths. It also differs in usually having well-developed anthers, and in being tetraploid. The two species are geographically isolated and ecologically distinct. Plants from the Columbia River Gorge in Oregon, including P. xmultnomae Piper, that approach P. tenerrima (p. 588) are presumed to be derived from hybridization between P. nervosa (see previous) and P. secunda (p. 586).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 24, p. 538. FNA vol. 24, p. 545.
Parent taxa Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Homalopoa Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Madropoa > subsect. Poa
Sibling taxa
P. abbreviata, P. alpina, P. alsodes, P. ammophila, P. annua, P. arachnifera, P. arctica, P. arida, P. arnowiae, P. atropurpurea, P. autumnalis, P. bigelovii, P. bolanderi, P. bulbosa, P. chaixii, P. chambersii, P. chapmaniana, P. compressa, P. confinis, P. curtifolia, P. cusickii, P. cuspidata, P. diaboli, P. douglasii, P. eminens, P. fendleriana, P. glauca, P. hartzii, P. howellii, P. infirma, P. interior, P. keckii, P. kelloggii, P. laxa, P. laxa × glauca, P. laxiflora, P. leibergii, P. leptocoma, P. lettermanii, P. macrantha, P. macrocalyx, P. marcida, P. napensis, P. nemoralis, P. nervosa, P. occidentalis, P. paludigena, P. palustris, P. paucispicula, P. piperi, P. porsildii, P. pratensis, P. pringlei, P. pseudoabbreviata, P. rhizomata, P. saltuensis, P. secunda, P. sierrae, P. stebbinsii, P. stenantha, P. strictiramea, P. sublanata, P. suksdorfii, P. supina, P. sylvestris, P. tenerrima, P. tracyi, P. trivialis, P. unilateralis, P. wheeleri, P. wolfii, P. ×gaspensis, P. ×limosa, P. ×nematophylla
P. abbreviata, P. alpina, P. alsodes, P. ammophila, P. annua, P. arachnifera, P. arctica, P. arida, P. arnowiae, P. atropurpurea, P. autumnalis, P. bigelovii, P. bolanderi, P. bulbosa, P. chaixii, P. chambersii, P. chapmaniana, P. compressa, P. confinis, P. curtifolia, P. cusickii, P. cuspidata, P. diaboli, P. douglasii, P. eminens, P. fendleriana, P. glauca, P. hartzii, P. howellii, P. infirma, P. interior, P. keckii, P. kelloggii, P. laxa, P. laxa × glauca, P. laxiflora, P. leibergii, P. leptocoma, P. lettermanii, P. macrantha, P. macrocalyx, P. marcida, P. napensis, P. nemoralis, P. occidentalis, P. paludigena, P. palustris, P. paucispicula, P. piperi, P. porsildii, P. pratensis, P. pringlei, P. pseudoabbreviata, P. reflexa, P. rhizomata, P. saltuensis, P. secunda, P. sierrae, P. stebbinsii, P. stenantha, P. strictiramea, P. sublanata, P. suksdorfii, P. supina, P. sylvestris, P. tenerrima, P. tracyi, P. trivialis, P. unilateralis, P. wheeleri, P. wolfii, P. ×gaspensis, P. ×limosa, P. ×nematophylla
Name authority Vasey & Scribn. (Hook.) Vasey
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