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nodding bluegrass

large-glume bluegrass

Habit Plants perennial, short-lived; densely tufted, tuft bases narrow or not, not stoloniferous, not rhizomatous. Plants perennial; green or strongly purplish; rhizomatous, rhizomes sometimes poorly developed, densely to loosely tufted.
Culms

10-60 cm.

(20)30-120 cm, usually stout, erect or decumbent, not branching above the base, terete or weakly compressed;

nodes terete, 1-2 exserted.

Sheaths

closed for 1/3 – 2/3 their length, terete, smooth;

ligules 1.5-3.5 mm, smooth or sparsely scabrous;

blades 1.5-4 mm wide, flat, thin, soft, apices broadly prow-shaped.

closed for 1/4 - 3/5 their length, terete to compressed, usually distinctly keeled, glabrous, sparsely to moderately scabrous, bases of basal sheaths glabrous, distal sheath lengths (0.8)1-2(2.5) times blade lengths;

collars smooth, glabrous;

ligules 2-5(6) mm, usually smooth, ligules of proximal leaves sometimes scabrous, obtuse, those of the distal leaves acute;

blades 4-12(18) cm long, (2)3-7 mm wide, flat or loosely folded, glabrous, both surfaces smooth or the adaxial surfaces sparsely scabrous, apices broadly prow-shaped, cauline blades subequal, flag leaf blades 3-10 cm.

Basal branching

mixed intra- and extravaginal.

all or mainly extravaginal.

Panicles

4-15 cm, nodding, open, with numerous spikelets and 1-2 branches per node;

branches (2)3-7 cm, spreading to reflexed, lower branches usually reflexed, flexuous, usually terete, smooth or sparsely scabrous, with (3)6-18 spikelets.

(4)7-15(20) cm, lax, loosely contracted or open, moderately congested, with (15)20-100 spikelets and 2-4(5) branches per node;

branches 3-8 cm, ascending or eventually spreading, flexuous, terete or weakly angled, usually sparsely scabridulous or scabrous, infrequently moderately densely scabrous, with (3)5-15(30) spikelets moderately crowded in the distal 1/2.

Spikelets

4-6 mm, lanceolate to broadly lanceolate, usually partly to wholly purplish, with 3-5 florets;

rachilla internodes shorter than 1 mm, smooth.

(5)6-9(12) mm, lengths to 3 times widths, laterally compressed, sometimes bulbiferous;

florets 2-5, infrequently bulb-forming;

rachilla internodes smooth or sparsely hispidulous.

Glumes

narrowly to broadly lanceolate, distinctly keeled, keels smooth or nearly so;

lower glumes 1-veined;

upper glumes shorter than or subequal to the lowest lemmas;

calluses webbed;

lemmas 2-3.5 mm, lanceolate, partly purple to fairly strongly purple, distinctly keeled, keels and marginal veins short- to long-villous, keels hairy for 2h-ls their length, lateral veins usually sparsely softly puberulent at least on 1 side, lateral veins obscure to moderately prominent, intercostal regions smooth, minutely bumpy, glabrous, apices acute, slightly bronze-colored or not;

palea keels scabrous, usually softly puberulent at midlength;

anthers 0.6-1 mm.

subequal in length and width, usually nearly equaling the adjacent lemmas, lanceolate to broadly lanceolate, distinctly keeled, keels sparsely to moderately scabrous distally, lateral veins prominent;

lower glumes (4)4.5-7 mm, 3-veined: upper glumes (4)4.5-7(8) mm, 3(5)-veined;

calluses dorsally webbed, hairs 1/3 – 2/3 times the lemma length, copious;

lemmas (4)5-8 mm, lanceolate to broadly lanceolate, green or purple, distinctly keeled, keels and marginal veins long-villous, lateral veins prominent, usually softly puberulent to short-villous, intercostal regions sometimes hispidulous, infrequently softly puberulent or short-villous proximally and hispidulous in the central 1/3, sometimes merely muriculate proximally and scabrous distally, sometimes densely scabrous throughout, rarely glabrous and nearly smooth proximally and sparsely scabrous distally, margins broadly hyaline, glabrous, apices acute;

palea keels scabrous, sometimes softly puberulent at midlength, intercostal regions broad, distinctly hispidulous or softly puberulent;

anthers 1.5-2.5 mm.

2n

= 28.

= 42, 43, 44, 45, ca. 46, 49, 56, ca. 58, 63, ca. 64, ca. 65, 66, 67, 68, 69, 70, ca. 71, ca. 72, ca. 73, 74, 75, 76, 77, 78, 80, ca. 82, ca. 84, ca. 87, ca. 100.

Poa reflexa

Poa macrocalyx

Distribution
from FNA
AZ; CO; ID; MT; NM; NV; OR; UT; WY
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; BC
[BONAP county map]
Discussion

Poa reflexa grows in subalpine forests, meadows, and low alpine habitats, primarily in the central and southern Rocky Mountains. It usually grows on drier and more disturbed sites, and appears shorter-lived, than the frequently sympatric or parapatric P. leptocoma (p. 573), from which it differs in usually having hairs on the palea keels and lateral veins of the lemmas, and smooth panicle branches. In addition, P. reflexa is tetraploid, whereas P. leptocoma is hexaploid. Poa reflexa may resemble small plants of P. occidentalis (see previous) in habit.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Poa macrocalyx grows mainly in coastal areas of boreal Alaska, and from the eastern coast of Russia to northern Japan. Bulbiferous plants are occasionally found. Poa macrocalyx resembles an exaggeratedly robust P. pratensis (p. 522), with large spikelets and lemmas, proportionally longer glumes, and paleas that are generally hispidulous between the veins and palea keels. It is cytologically and morphologically complex, and is sometimes difficult to distinguish from P. arctica subsp. lanata (see below). Poa norbergii Hulten may belong to this species; alternatively, it may be a hybrid between P. glauca (p. 576) and P. arctica (p. 576) or P. macrocalyx.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 24, p. 538. FNA vol. 24, p. 527.
Parent taxa Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Homalopoa Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Poa
Sibling taxa
P. abbreviata, P. alpina, P. alsodes, P. ammophila, P. annua, P. arachnifera, P. arctica, P. arida, P. arnowiae, P. atropurpurea, P. autumnalis, P. bigelovii, P. bolanderi, P. bulbosa, P. chaixii, P. chambersii, P. chapmaniana, P. compressa, P. confinis, P. curtifolia, P. cusickii, P. cuspidata, P. diaboli, P. douglasii, P. eminens, P. fendleriana, P. glauca, P. hartzii, P. howellii, P. infirma, P. interior, P. keckii, P. kelloggii, P. laxa, P. laxa × glauca, P. laxiflora, P. leibergii, P. leptocoma, P. lettermanii, P. macrantha, P. macrocalyx, P. marcida, P. napensis, P. nemoralis, P. nervosa, P. occidentalis, P. paludigena, P. palustris, P. paucispicula, P. piperi, P. porsildii, P. pratensis, P. pringlei, P. pseudoabbreviata, P. rhizomata, P. saltuensis, P. secunda, P. sierrae, P. stebbinsii, P. stenantha, P. strictiramea, P. sublanata, P. suksdorfii, P. supina, P. sylvestris, P. tenerrima, P. tracyi, P. trivialis, P. unilateralis, P. wheeleri, P. wolfii, P. ×gaspensis, P. ×limosa, P. ×nematophylla
P. abbreviata, P. alpina, P. alsodes, P. ammophila, P. annua, P. arachnifera, P. arctica, P. arida, P. arnowiae, P. atropurpurea, P. autumnalis, P. bigelovii, P. bolanderi, P. bulbosa, P. chaixii, P. chambersii, P. chapmaniana, P. compressa, P. confinis, P. curtifolia, P. cusickii, P. cuspidata, P. diaboli, P. douglasii, P. eminens, P. fendleriana, P. glauca, P. hartzii, P. howellii, P. infirma, P. interior, P. keckii, P. kelloggii, P. laxa, P. laxa × glauca, P. laxiflora, P. leibergii, P. leptocoma, P. lettermanii, P. macrantha, P. marcida, P. napensis, P. nemoralis, P. nervosa, P. occidentalis, P. paludigena, P. palustris, P. paucispicula, P. piperi, P. porsildii, P. pratensis, P. pringlei, P. pseudoabbreviata, P. reflexa, P. rhizomata, P. saltuensis, P. secunda, P. sierrae, P. stebbinsii, P. stenantha, P. strictiramea, P. sublanata, P. suksdorfii, P. supina, P. sylvestris, P. tenerrima, P. tracyi, P. trivialis, P. unilateralis, P. wheeleri, P. wolfii, P. ×gaspensis, P. ×limosa, P. ×nematophylla
Synonyms P. turneri, P. ×norbergii, P. hispidula
Name authority Vasey & Scribn. Trautv. & CA. Mey.
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