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nodding bluegrass

Douglas' bluegrass, sand dune blue grass

Habit Plants perennial, short-lived; densely tufted, tuft bases narrow or not, not stoloniferous, not rhizomatous. Plants perennial; loosely tufted, rhizomatous and stoloniferous, rhizomes and stolons to 1 m. Basal branching mainly intravaginal, some extravaginal.
Culms

10-60 cm.

(5)10-30 cm tall, 1.2-1.5 mm thick, bases decumbent, terete or weakly compressed, hispidulous beneath the panicles;

nodes terete, 0(1) exserted.

Sheaths

closed for 1/3 – 2/3 their length, terete, smooth;

ligules 1.5-3.5 mm, smooth or sparsely scabrous;

blades 1.5-4 mm wide, flat, thin, soft, apices broadly prow-shaped.

closed for about 1/2 their length, terete, smooth or sparsely to moderately retrorsely scabrous near the collars, glabrous, bases of basal sheaths glabrous, distal sheath lengths 0.9-3.5 times blade lengths;

collars sparsely to moderately retrorsely scabrous, glabrous;

ligules 1-2 mm, scabrous, truncate to obtuse, ciliolate;

innovation blades to 30 cm long, adaxial surfaces moderately to densely scabrous or hispidulous on and between the veins;

cauline blades subequal in length, 1-2 mm wide, involute, moderately thick, moderately firm, arcuate, abaxial surfaces smooth or sparsely scabrous, adaxial surfaces moderately to densely scabrous or hispidulous on and between the veins, apices narrowly prow-shaped, flag leaf blades 1-9 cm.

Basal branching

mixed intra- and extravaginal.

Panicles

4-15 cm, nodding, open, with numerous spikelets and 1-2 branches per node;

branches (2)3-7 cm, spreading to reflexed, lower branches usually reflexed, flexuous, usually terete, smooth or sparsely scabrous, with (3)6-18 spikelets.

1.5-6 cm, erect, compact, ovoid, contracted, infrequently interrupted, congested, with 15-50 spikelets;

nodes with 1-2 branches, internodes densely hispidulous;

branches 0.5-2 cm, erect, stiff, terete to weakly angled, densely hispidulous, with 1-5 spikelets.

Spikelets

4-6 mm, lanceolate to broadly lanceolate, usually partly to wholly purplish, with 3-5 florets;

rachilla internodes shorter than 1 mm, smooth.

7-12 mm, lengths to 3 times widths, laterally compressed, not sexually dimorphic;

florets 3-6;

rachilla internodes usually shorter than 0.5 mm, smooth, glabrous.

Glumes

narrowly to broadly lanceolate, distinctly keeled, keels smooth or nearly so;

lower glumes 1-veined;

upper glumes shorter than or subequal to the lowest lemmas;

calluses webbed;

lemmas 2-3.5 mm, lanceolate, partly purple to fairly strongly purple, distinctly keeled, keels and marginal veins short- to long-villous, keels hairy for 2h-ls their length, lateral veins usually sparsely softly puberulent at least on 1 side, lateral veins obscure to moderately prominent, intercostal regions smooth, minutely bumpy, glabrous, apices acute, slightly bronze-colored or not;

palea keels scabrous, usually softly puberulent at midlength;

anthers 0.6-1 mm.

broadly lanceolate, 1/2 as long as to subequal to the adjacent lemmas, distinctly keeled;

lower glumes 3-veined;

upper glumes 4-4.5(7+) mm, 3-veined;

calluses usually with a crown of hairs, sometimes glabrous or diffusely webbed;

lemmas 5-7.5 mm, lanceolate, 5-veined, distinctly keeled, keels, marginal veins, and sometimes the lateral veins short- to long-villous or softly puberulent, rarely glabrous, lateral veins moderately prominent, intercostal regions smooth, glabrous, margins glabrous, apices acute;

palea keels scabrous to pectinate-ciliate, intercostal regions glabrous;

anthers vestigial (0.1-0.2 mm) or (2)2.5-3.5 (4) mm.

2n

= 28.

= 28.

Poa reflexa

Poa douglasii

Distribution
from FNA
AZ; CO; ID; MT; NM; NV; OR; UT; WY
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
Discussion

Poa reflexa grows in subalpine forests, meadows, and low alpine habitats, primarily in the central and southern Rocky Mountains. It usually grows on drier and more disturbed sites, and appears shorter-lived, than the frequently sympatric or parapatric P. leptocoma (p. 573), from which it differs in usually having hairs on the palea keels and lateral veins of the lemmas, and smooth panicle branches. In addition, P. reflexa is tetraploid, whereas P. leptocoma is hexaploid. Poa reflexa may resemble small plants of P. occidentalis (see previous) in habit.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Poa douglasii is a dioecious endemic that grows on coastal sand dunes in California, a habitat that is being invaded by exotic species. It is rare north of Mendocino. Its hairy rachises distinguish P. douglasii from all other species of Poa in the Flora region. It differs from P. macrantha (see next), which occupies similar habitats, in this and in its usually longer glumes and lemmas.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 24, p. 538. FNA vol. 24, p. 551.
Parent taxa Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Homalopoa Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Madropoa > subsect. Madropoa
Sibling taxa
P. abbreviata, P. alpina, P. alsodes, P. ammophila, P. annua, P. arachnifera, P. arctica, P. arida, P. arnowiae, P. atropurpurea, P. autumnalis, P. bigelovii, P. bolanderi, P. bulbosa, P. chaixii, P. chambersii, P. chapmaniana, P. compressa, P. confinis, P. curtifolia, P. cusickii, P. cuspidata, P. diaboli, P. douglasii, P. eminens, P. fendleriana, P. glauca, P. hartzii, P. howellii, P. infirma, P. interior, P. keckii, P. kelloggii, P. laxa, P. laxa × glauca, P. laxiflora, P. leibergii, P. leptocoma, P. lettermanii, P. macrantha, P. macrocalyx, P. marcida, P. napensis, P. nemoralis, P. nervosa, P. occidentalis, P. paludigena, P. palustris, P. paucispicula, P. piperi, P. porsildii, P. pratensis, P. pringlei, P. pseudoabbreviata, P. rhizomata, P. saltuensis, P. secunda, P. sierrae, P. stebbinsii, P. stenantha, P. strictiramea, P. sublanata, P. suksdorfii, P. supina, P. sylvestris, P. tenerrima, P. tracyi, P. trivialis, P. unilateralis, P. wheeleri, P. wolfii, P. ×gaspensis, P. ×limosa, P. ×nematophylla
P. abbreviata, P. alpina, P. alsodes, P. ammophila, P. annua, P. arachnifera, P. arctica, P. arida, P. arnowiae, P. atropurpurea, P. autumnalis, P. bigelovii, P. bolanderi, P. bulbosa, P. chaixii, P. chambersii, P. chapmaniana, P. compressa, P. confinis, P. curtifolia, P. cusickii, P. cuspidata, P. diaboli, P. eminens, P. fendleriana, P. glauca, P. hartzii, P. howellii, P. infirma, P. interior, P. keckii, P. kelloggii, P. laxa, P. laxa × glauca, P. laxiflora, P. leibergii, P. leptocoma, P. lettermanii, P. macrantha, P. macrocalyx, P. marcida, P. napensis, P. nemoralis, P. nervosa, P. occidentalis, P. paludigena, P. palustris, P. paucispicula, P. piperi, P. porsildii, P. pratensis, P. pringlei, P. pseudoabbreviata, P. reflexa, P. rhizomata, P. saltuensis, P. secunda, P. sierrae, P. stebbinsii, P. stenantha, P. strictiramea, P. sublanata, P. suksdorfii, P. supina, P. sylvestris, P. tenerrima, P. tracyi, P. trivialis, P. unilateralis, P. wheeleri, P. wolfii, P. ×gaspensis, P. ×limosa, P. ×nematophylla
Name authority Vasey & Scribn. Nees
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