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Howell's blue grass

nodding bluegrass

Habit Plants usually annual, rarely longer-lived; densely tufted, tuft bases narrow; not stoloniferous, not rhizomatous. Plants perennial, short-lived; densely tufted, tuft bases narrow or not, not stoloniferous, not rhizomatous.
Culms

(10) 25-80(120) cm tall, 0.4-1.75 mm thick, usually erect;

nodes terete, usually 1-2 exserted.

10-60 cm.

Sheaths

closed for 1/2 - 7/8 their length, usually weakly compressed and keeled, usually scabrous, rarely smooth;

ligules 1.5-5(10) mm, smooth or scabrous, acute;

blades 1-7(10) mm wide, flat, lax, soft, finely scabrous, apices narrowly prow-shaped, cauline blades 2-10 cm.

closed for 1/3 – 2/3 their length, terete, smooth;

ligules 1.5-3.5 mm, smooth or sparsely scabrous;

blades 1.5-4 mm wide, flat, thin, soft, apices broadly prow-shaped.

Basal branching

intravaginal.

mixed intra- and extravaginal.

Panicles

10-25(30) cm, erect, eventually open, with (1)3-5(7) branches per node;

branches eventually spreading or reflexed, fairly straight, angled, angles usually moderately to densely scabrous, rarely sparsely scabrous.

4-15 cm, nodding, open, with numerous spikelets and 1-2 branches per node;

branches (2)3-7 cm, spreading to reflexed, lower branches usually reflexed, flexuous, usually terete, smooth or sparsely scabrous, with (3)6-18 spikelets.

Spikelets

(2)4-6 mm, laterally compressed, with 2-5 florets;

rachilla internodes about 1 mm, smooth, usually softly puberulent, infrequently glabrous.

4-6 mm, lanceolate to broadly lanceolate, usually partly to wholly purplish, with 3-5 florets;

rachilla internodes shorter than 1 mm, smooth.

Glumes

slightly unequal, lanceolate, distinctly keeled, keels and sometimes the lateral veins sparsely to moderately scabrous;

lower glumes 1-3-veined;

upper glumes shorter than or subequal to the lowest lemmas;

calluses of some or all florets sparsely webbed;

lemmas 2.5-3.5 mm, lanceolate to narrowly lanceolate, distinctly keeled, crisply puberulent proximally, hairs evenly distributed, finely scabrous distally, lateral veins obscure to prominent, margins narrowly hyaline, glabrous, apices narrowly acute, infrequently anthocyanic;

palea keels sparsely scabrous, glabrous or softly puberulent at midlength, intercostal regions usually softly puberulent;

anthers 3, 0.2-1 mm.

narrowly to broadly lanceolate, distinctly keeled, keels smooth or nearly so;

lower glumes 1-veined;

upper glumes shorter than or subequal to the lowest lemmas;

calluses webbed;

lemmas 2-3.5 mm, lanceolate, partly purple to fairly strongly purple, distinctly keeled, keels and marginal veins short- to long-villous, keels hairy for 2h-ls their length, lateral veins usually sparsely softly puberulent at least on 1 side, lateral veins obscure to moderately prominent, intercostal regions smooth, minutely bumpy, glabrous, apices acute, slightly bronze-colored or not;

palea keels scabrous, usually softly puberulent at midlength;

anthers 0.6-1 mm.

2n

= unknown.

= 28.

Poa howellii

Poa reflexa

Distribution
from FNA
CA; OR; WA; BC
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CO; ID; MT; NM; NV; OR; UT; WY
[WildflowerSearch map]
[BONAP county map]
Discussion

Poa howellii grows primarily on rocky banks and wooded slopes, from the coastal ranges of southern British Columbia to southern California. It differs from P. bolanderi (see above) in having puberulent, rather than smooth or scabrous, lemmas, and in growing at lower elevations, mostly from near sea level to 1000 m.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Poa reflexa grows in subalpine forests, meadows, and low alpine habitats, primarily in the central and southern Rocky Mountains. It usually grows on drier and more disturbed sites, and appears shorter-lived, than the frequently sympatric or parapatric P. leptocoma (p. 573), from which it differs in usually having hairs on the palea keels and lateral veins of the lemmas, and smooth panicle branches. In addition, P. reflexa is tetraploid, whereas P. leptocoma is hexaploid. Poa reflexa may resemble small plants of P. occidentalis (see previous) in habit.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 24, p. 534. FNA vol. 24, p. 538.
Parent taxa Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Homalopoa Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Homalopoa
Sibling taxa
P. abbreviata, P. alpina, P. alsodes, P. ammophila, P. annua, P. arachnifera, P. arctica, P. arida, P. arnowiae, P. atropurpurea, P. autumnalis, P. bigelovii, P. bolanderi, P. bulbosa, P. chaixii, P. chambersii, P. chapmaniana, P. compressa, P. confinis, P. curtifolia, P. cusickii, P. cuspidata, P. diaboli, P. douglasii, P. eminens, P. fendleriana, P. glauca, P. hartzii, P. infirma, P. interior, P. keckii, P. kelloggii, P. laxa, P. laxa × glauca, P. laxiflora, P. leibergii, P. leptocoma, P. lettermanii, P. macrantha, P. macrocalyx, P. marcida, P. napensis, P. nemoralis, P. nervosa, P. occidentalis, P. paludigena, P. palustris, P. paucispicula, P. piperi, P. porsildii, P. pratensis, P. pringlei, P. pseudoabbreviata, P. reflexa, P. rhizomata, P. saltuensis, P. secunda, P. sierrae, P. stebbinsii, P. stenantha, P. strictiramea, P. sublanata, P. suksdorfii, P. supina, P. sylvestris, P. tenerrima, P. tracyi, P. trivialis, P. unilateralis, P. wheeleri, P. wolfii, P. ×gaspensis, P. ×limosa, P. ×nematophylla
P. abbreviata, P. alpina, P. alsodes, P. ammophila, P. annua, P. arachnifera, P. arctica, P. arida, P. arnowiae, P. atropurpurea, P. autumnalis, P. bigelovii, P. bolanderi, P. bulbosa, P. chaixii, P. chambersii, P. chapmaniana, P. compressa, P. confinis, P. curtifolia, P. cusickii, P. cuspidata, P. diaboli, P. douglasii, P. eminens, P. fendleriana, P. glauca, P. hartzii, P. howellii, P. infirma, P. interior, P. keckii, P. kelloggii, P. laxa, P. laxa × glauca, P. laxiflora, P. leibergii, P. leptocoma, P. lettermanii, P. macrantha, P. macrocalyx, P. marcida, P. napensis, P. nemoralis, P. nervosa, P. occidentalis, P. paludigena, P. palustris, P. paucispicula, P. piperi, P. porsildii, P. pratensis, P. pringlei, P. pseudoabbreviata, P. rhizomata, P. saltuensis, P. secunda, P. sierrae, P. stebbinsii, P. stenantha, P. strictiramea, P. sublanata, P. suksdorfii, P. supina, P. sylvestris, P. tenerrima, P. tracyi, P. trivialis, P. unilateralis, P. wheeleri, P. wolfii, P. ×gaspensis, P. ×limosa, P. ×nematophylla
Synonyms P. bolanderi var. howellii
Name authority Vasey & Scribn. Vasey & Scribn.
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