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Howell's blue grass

Eurasian woodland bluegrass, forest blue grass, interior bluegrass (ssp. interior), wood blue grass, woodland bluegrass, woods bluegrass (ssp. nemoralis)

Habit Plants usually annual, rarely longer-lived; densely tufted, tuft bases narrow; not stoloniferous, not rhizomatous. Plants perennial; green or glaucous; densely tufted, not stoloniferous, not rhizomatous.
Culms

(10) 25-80(120) cm tall, 0.4-1.75 mm thick, usually erect;

nodes terete, usually 1-2 exserted.

30-80 cm, mostly erect, smooth below the panicles;

nodes slightly compressed, 2-5 exserted, top node at 1/2 - 3/4 the culm length.

Sheaths

closed for 1/2 - 7/8 their length, usually weakly compressed and keeled, usually scabrous, rarely smooth;

ligules 1.5-5(10) mm, smooth or scabrous, acute;

blades 1-7(10) mm wide, flat, lax, soft, finely scabrous, apices narrowly prow-shaped, cauline blades 2-10 cm.

closed for 1/10 – 1/5 their length, terete, bases of basal sheaths glabrous, distal sheath lengths 0.45-1 (1.1) times blade lengths;

ligules 0.2-0.8(1) mm, sparsely to densely scabrous, apices truncate, minutely ciliolate;

blades 0.8-3 mm wide, mostly flat, appressed, abruptly ascending to spreading, straight or somewhat lax, apices narrowly prow-shaped.

Basal branching

intravaginal.

all or mostly extravaginal.

Panicles

10-25(30) cm, erect, eventually open, with (1)3-5(7) branches per node;

branches eventually spreading or reflexed, fairly straight, angled, angles usually moderately to densely scabrous, rarely sparsely scabrous.

7-16(20) cm, lengths usually 2.5-4 times widths at maturity, usually erect, lax in shade forms, narrowly lanceoloid to ovoid, slightly to moderately congested;

nodes with 2-5 branches;

branches ascending to widely spreading, fairly straight, slender to moderately stout, angled, angles moderately to densely scabrous.

Spikelets

(2)4-6 mm, laterally compressed, with 2-5 florets;

rachilla internodes about 1 mm, smooth, usually softly puberulent, infrequently glabrous.

3-8 mm, lengths 2.5-3.5 times widths, narrowly lanceolate to lanceolate, laterally compressed, usually not glaucous;

florets (1)2-5;

rachilla internodes usually shorter than 1 mm, smooth, muriculate, or scabrous, usually puberulent, infrequently hispidulous or glabrous.

Glumes

slightly unequal, lanceolate, distinctly keeled, keels and sometimes the lateral veins sparsely to moderately scabrous;

lower glumes 1-3-veined;

upper glumes shorter than or subequal to the lowest lemmas;

calluses of some or all florets sparsely webbed;

lemmas 2.5-3.5 mm, lanceolate to narrowly lanceolate, distinctly keeled, crisply puberulent proximally, hairs evenly distributed, finely scabrous distally, lateral veins obscure to prominent, margins narrowly hyaline, glabrous, apices narrowly acute, infrequently anthocyanic;

palea keels sparsely scabrous, glabrous or softly puberulent at midlength, intercostal regions usually softly puberulent;

anthers 3, 0.2-1 mm.

subulate to narrowly lanceolate, distinctly keeled, keels smooth or sparsely scabrous, apices sharply acute to acuminate;

lower glumes 3-veined, long-tapered to a slender point, lengths 6.4-11 times widths;

upper glumes shorter than or subequal to the lowest lemmas;

calluses webbed, hairs sparse, often short;

lemmas 2.4-4 mm, proximal lemma widths less than 1/5 times lengths, narrowly lanceolate to lanceolate, distinctly keeled, keels and marginal veins short-villous, lateral veins glabrous, obscure, intercostal regions smooth or muriculate, glabrous, margins glabrous, apices acute, usually partially bronze-colored;

palea keels scabrous, intercostal regions glabrous;

anthers 0.8-1.9 mm.

2n

= unknown.

= 28, 35, 42, 48, 50, 56.

Poa howellii

Poa nemoralis

Distribution
from FNA
CA; OR; WA; BC
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; AZ; CA; CO; CT; DE; IA; ID; IL; IN; KY; MA; MD; ME; MI; MN; MO; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OR; PA; RI; SD; TN; TX; UT; VA; VT; WA; WI; WY; AB; BC; MB; NB; NL; NS; NT; ON; QC; SK; YT; Greenland
[WildflowerSearch map]
[BONAP county map]
Discussion

Poa howellii grows primarily on rocky banks and wooded slopes, from the coastal ranges of southern British Columbia to southern California. It differs from P. bolanderi (see above) in having puberulent, rather than smooth or scabrous, lemmas, and in growing at lower elevations, mostly from near sea level to 1000 m.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Introduced from northern Eurasia, Poa nemoralis is established primarily at low elevations in deciduous and mixed conifer/deciduous forests. It is now common in southeastern Canada and the northeastern United States, and is spreading in the west. It can be distinguished from P. glauca (p. 576) and P. interior (see next) by its consistently short ligules, high top culm node, relatively long flag leaf blades, and narrow glumes and lemmas. It is usually hexaploid.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 24, p. 534. FNA vol. 24, p. 574.
Parent taxa Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Homalopoa Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Stenopoa
Sibling taxa
P. abbreviata, P. alpina, P. alsodes, P. ammophila, P. annua, P. arachnifera, P. arctica, P. arida, P. arnowiae, P. atropurpurea, P. autumnalis, P. bigelovii, P. bolanderi, P. bulbosa, P. chaixii, P. chambersii, P. chapmaniana, P. compressa, P. confinis, P. curtifolia, P. cusickii, P. cuspidata, P. diaboli, P. douglasii, P. eminens, P. fendleriana, P. glauca, P. hartzii, P. infirma, P. interior, P. keckii, P. kelloggii, P. laxa, P. laxa × glauca, P. laxiflora, P. leibergii, P. leptocoma, P. lettermanii, P. macrantha, P. macrocalyx, P. marcida, P. napensis, P. nemoralis, P. nervosa, P. occidentalis, P. paludigena, P. palustris, P. paucispicula, P. piperi, P. porsildii, P. pratensis, P. pringlei, P. pseudoabbreviata, P. reflexa, P. rhizomata, P. saltuensis, P. secunda, P. sierrae, P. stebbinsii, P. stenantha, P. strictiramea, P. sublanata, P. suksdorfii, P. supina, P. sylvestris, P. tenerrima, P. tracyi, P. trivialis, P. unilateralis, P. wheeleri, P. wolfii, P. ×gaspensis, P. ×limosa, P. ×nematophylla
P. abbreviata, P. alpina, P. alsodes, P. ammophila, P. annua, P. arachnifera, P. arctica, P. arida, P. arnowiae, P. atropurpurea, P. autumnalis, P. bigelovii, P. bolanderi, P. bulbosa, P. chaixii, P. chambersii, P. chapmaniana, P. compressa, P. confinis, P. curtifolia, P. cusickii, P. cuspidata, P. diaboli, P. douglasii, P. eminens, P. fendleriana, P. glauca, P. hartzii, P. howellii, P. infirma, P. interior, P. keckii, P. kelloggii, P. laxa, P. laxa × glauca, P. laxiflora, P. leibergii, P. leptocoma, P. lettermanii, P. macrantha, P. macrocalyx, P. marcida, P. napensis, P. nervosa, P. occidentalis, P. paludigena, P. palustris, P. paucispicula, P. piperi, P. porsildii, P. pratensis, P. pringlei, P. pseudoabbreviata, P. reflexa, P. rhizomata, P. saltuensis, P. secunda, P. sierrae, P. stebbinsii, P. stenantha, P. strictiramea, P. sublanata, P. suksdorfii, P. supina, P. sylvestris, P. tenerrima, P. tracyi, P. trivialis, P. unilateralis, P. wheeleri, P. wolfii, P. ×gaspensis, P. ×limosa, P. ×nematophylla
Synonyms P. bolanderi var. howellii
Name authority Vasey & Scribn. L.
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