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Howell's blue grass

banff bluegrass, glaucous bluegrass, lax bluegrass, Mt. Washington bluegrass, wavy blue grass

Habit Plants usually annual, rarely longer-lived; densely tufted, tuft bases narrow; not stoloniferous, not rhizomatous. Plants perennial; not or only slightly glaucous; densely tufted, not stoloniferous, not rhizomatous.
Culms

(10) 25-80(120) cm tall, 0.4-1.75 mm thick, usually erect;

nodes terete, usually 1-2 exserted.

8-35 cm tall, 0.5-0.9 mm thick, ascending to erect, slender;

nodes terete, 0(1) exserted.

Sheaths

closed for 1/2 - 7/8 their length, usually weakly compressed and keeled, usually scabrous, rarely smooth;

ligules 1.5-5(10) mm, smooth or scabrous, acute;

blades 1-7(10) mm wide, flat, lax, soft, finely scabrous, apices narrowly prow-shaped, cauline blades 2-10 cm.

closed for 1/5-1/3 their length, terete, smooth, glabrous, bases of basal sheaths glabrous;

collars smooth or scabrous, glabrous;

ligules 2-4 mm, smooth, apices acute, often lacerate;

innovation blades similar to the cauline blades;

cauline blades 1-2(3) mm wide, flat, thin, soft, smooth, narrowly prow-tipped, blades not strongly graduated or reduced upwards.

Basal branching

intravaginal.

mixed, mainly extravaginal or mainly pseudointravaginal, sometimes intravaginal.

Panicles

10-25(30) cm, erect, eventually open, with (1)3-5(7) branches per node;

branches eventually spreading or reflexed, fairly straight, angled, angles usually moderately to densely scabrous, rarely sparsely scabrous.

2-8 cm, slightly lax, usually loosely contracted and sparse, infrequently contracted and dense;

nodes with 1-3(5) branches;

branches 1-3(4) cm, usually ascending or weakly spreading, infrequently erect, fairly straight or flexuous, slender, sulcate or angled, smooth or the angles sparsely scabrous, with 1-8 spikelets.

Spikelets

(2)4-6 mm, laterally compressed, with 2-5 florets;

rachilla internodes about 1 mm, smooth, usually softly puberulent, infrequently glabrous.

4-6 mm, lengths to 3 times widths, laterally compressed;

florets 2-5;

rachilla internodes shorter than 1 mm, smooth, glabrous.

Glumes

slightly unequal, lanceolate, distinctly keeled, keels and sometimes the lateral veins sparsely to moderately scabrous;

lower glumes 1-3-veined;

upper glumes shorter than or subequal to the lowest lemmas;

calluses of some or all florets sparsely webbed;

lemmas 2.5-3.5 mm, lanceolate to narrowly lanceolate, distinctly keeled, crisply puberulent proximally, hairs evenly distributed, finely scabrous distally, lateral veins obscure to prominent, margins narrowly hyaline, glabrous, apices narrowly acute, infrequently anthocyanic;

palea keels sparsely scabrous, glabrous or softly puberulent at midlength, intercostal regions usually softly puberulent;

anthers 3, 0.2-1 mm.

nearly equaling or slightly longer than the adjacent lemmas, lanceolate to broadly lanceolate, thin, distinctly keeled, keels smooth or sparsely scabrous;

lower glumes 1-3-veined;

upper glumes shorter than or subequal to the lowest lemmas;

calluses glabrous or webbed, hairs usually shorter than 1/4 the lemma length, sparse;

lemmas 3-4.6 mm, lanceolate to broadly lanceolate, thin, distinctly keeled, keels and marginal veins short- to long-villous, lateral veins glabrous or sparsely softly puberulent, lateral veins obscure, intercostal regions glabrous, margins glabrous, apices acute;

paleas sparsely scabrous over the keels;

anthers (0.6)0.8-1.1(1.3) mm.

2n

= unknown.

= 28, 42, 84.

Poa howellii

Poa laxa

Distribution
from FNA
CA; OR; WA; BC
[WildflowerSearch map]
[BONAP county map]
from FNA
CO; ME; MT; NH; NY; OR; VT; WY; AB; BC; NL; QC
[BONAP county map]
Discussion

Poa howellii grows primarily on rocky banks and wooded slopes, from the coastal ranges of southern British Columbia to southern California. It differs from P. bolanderi (see above) in having puberulent, rather than smooth or scabrous, lemmas, and in growing at lower elevations, mostly from near sea level to 1000 m.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Poa laxa is a low arctic to high alpine amphiatlantic species. It has been treated as a series of separate species, but the differences seem relatively minor and incomplete. Its short anthers and smoother branches usually distinguish it from P. glauca (p. 576), with which it can hybridize to form P. laxa x glauca (p. 572).

Poa laxa has four subspecies, two of which are native to the Flora region; subsp. laxa grows in central Europe; and subsp. flexuosa (Sm.) Hyl. in northwestern Europe.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Innovations primarily extravaginal; panicle branches fairly straight; calluses glabrous
subsp. banffiana
1. Innovations primarily intravaginal; panicle branches flexuous, usually at least some florets having a webbed callus
subsp. fernaldiana
Source FNA vol. 24, p. 534. FNA vol. 24, p. 570.
Parent taxa Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Homalopoa Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Oreinos
Sibling taxa
P. abbreviata, P. alpina, P. alsodes, P. ammophila, P. annua, P. arachnifera, P. arctica, P. arida, P. arnowiae, P. atropurpurea, P. autumnalis, P. bigelovii, P. bolanderi, P. bulbosa, P. chaixii, P. chambersii, P. chapmaniana, P. compressa, P. confinis, P. curtifolia, P. cusickii, P. cuspidata, P. diaboli, P. douglasii, P. eminens, P. fendleriana, P. glauca, P. hartzii, P. infirma, P. interior, P. keckii, P. kelloggii, P. laxa, P. laxa × glauca, P. laxiflora, P. leibergii, P. leptocoma, P. lettermanii, P. macrantha, P. macrocalyx, P. marcida, P. napensis, P. nemoralis, P. nervosa, P. occidentalis, P. paludigena, P. palustris, P. paucispicula, P. piperi, P. porsildii, P. pratensis, P. pringlei, P. pseudoabbreviata, P. reflexa, P. rhizomata, P. saltuensis, P. secunda, P. sierrae, P. stebbinsii, P. stenantha, P. strictiramea, P. sublanata, P. suksdorfii, P. supina, P. sylvestris, P. tenerrima, P. tracyi, P. trivialis, P. unilateralis, P. wheeleri, P. wolfii, P. ×gaspensis, P. ×limosa, P. ×nematophylla
P. abbreviata, P. alpina, P. alsodes, P. ammophila, P. annua, P. arachnifera, P. arctica, P. arida, P. arnowiae, P. atropurpurea, P. autumnalis, P. bigelovii, P. bolanderi, P. bulbosa, P. chaixii, P. chambersii, P. chapmaniana, P. compressa, P. confinis, P. curtifolia, P. cusickii, P. cuspidata, P. diaboli, P. douglasii, P. eminens, P. fendleriana, P. glauca, P. hartzii, P. howellii, P. infirma, P. interior, P. keckii, P. kelloggii, P. laxa × glauca, P. laxiflora, P. leibergii, P. leptocoma, P. lettermanii, P. macrantha, P. macrocalyx, P. marcida, P. napensis, P. nemoralis, P. nervosa, P. occidentalis, P. paludigena, P. palustris, P. paucispicula, P. piperi, P. porsildii, P. pratensis, P. pringlei, P. pseudoabbreviata, P. reflexa, P. rhizomata, P. saltuensis, P. secunda, P. sierrae, P. stebbinsii, P. stenantha, P. strictiramea, P. sublanata, P. suksdorfii, P. supina, P. sylvestris, P. tenerrima, P. tracyi, P. trivialis, P. unilateralis, P. wheeleri, P. wolfii, P. ×gaspensis, P. ×limosa, P. ×nematophylla
Subordinate taxa
P. laxa subsp. banffiana, P. laxa subsp. fernaldiana
Synonyms P. bolanderi var. howellii
Name authority Vasey & Scribn. Haenke
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