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Howell's blue grass

Diablo bluegrass, Diablo Canyon blue grass

Habit Plants usually annual, rarely longer-lived; densely tufted, tuft bases narrow; not stoloniferous, not rhizomatous. Plants perennial; loosely tufted, forming airy mounds to 30 cm across, shortly rhizomatous and stoloniferous.
Culms

(10) 25-80(120) cm tall, 0.4-1.75 mm thick, usually erect;

nodes terete, usually 1-2 exserted.

26-50 cm tall, 0.5-0.9 mm thick, bases decumbent or nearly erect, frequently branching above the base, terete or weakly compressed;

nodes terete, 1-2 exserted.

Sheaths

closed for 1/2 - 7/8 their length, usually weakly compressed and keeled, usually scabrous, rarely smooth;

ligules 1.5-5(10) mm, smooth or scabrous, acute;

blades 1-7(10) mm wide, flat, lax, soft, finely scabrous, apices narrowly prow-shaped, cauline blades 2-10 cm.

closed for 2/5 – 7/10 their length, weakly keeled, sparsely scabrous, glabrous, bases of basal sheaths glabrous, distal sheath lengths 0.6-2.4 times blade lengths;

collars scabrous or pubescent on the margins;

ligules (1)2-3 mm, moderately densely scabrous, truncate, obtuse, or acute, lacerate to entire;

innovation blades to 20 cm, adaxial surfaces sparsely scabrous, glabrous or hispidulous on and between the veins;

cauline blades 1.5-2.4 mm wide, folded or flat, thin, soft, abaxial surfaces smooth, veins prominent, keel and margins scabrous, adaxial surfaces moderately scabrous over the veins, sparsely scabrous between the veins, apices narrowly prow-shaped, flag leaf blades 2.9-8.6(11) cm.

Basal branching

intravaginal.

extra-, pseudo-, and intravaginal.

Panicles

10-25(30) cm, erect, eventually open, with (1)3-5(7) branches per node;

branches eventually spreading or reflexed, fairly straight, angled, angles usually moderately to densely scabrous, rarely sparsely scabrous.

(4)5.5-10.5(12.5) cm, erect, ovoid to broadly pyramidal, open, or eventually loosely contracted, sparse, with 10-40 spikelets;

nodes with 1-2 branches;

branches 2.1-4.5(7) cm, ascending, lax, angled, angles moderately to densely scabrous, less scabrous between the angles, with 1-9 spikelets.

Spikelets

(2)4-6 mm, laterally compressed, with 2-5 florets;

rachilla internodes about 1 mm, smooth, usually softly puberulent, infrequently glabrous.

5.3-9 mm, lengths to 3 times widths, laterally compressed, not sexually dimorphic;

florets (2)3-6(7);

rachilla internodes 1-1.3 mm, visible from the sides, usually sparsely to densely, coarsely scabrous, infrequently smooth.

Glumes

slightly unequal, lanceolate, distinctly keeled, keels and sometimes the lateral veins sparsely to moderately scabrous;

lower glumes 1-3-veined;

upper glumes shorter than or subequal to the lowest lemmas;

calluses of some or all florets sparsely webbed;

lemmas 2.5-3.5 mm, lanceolate to narrowly lanceolate, distinctly keeled, crisply puberulent proximally, hairs evenly distributed, finely scabrous distally, lateral veins obscure to prominent, margins narrowly hyaline, glabrous, apices narrowly acute, infrequently anthocyanic;

palea keels sparsely scabrous, glabrous or softly puberulent at midlength, intercostal regions usually softly puberulent;

anthers 3, 0.2-1 mm.

distinctly keeled;

lower glumes (2)2.7-3.8 mm, 3-veined, upper glumes (2.3)2.9-3.9 mm;

calluses diffusely webbed, hairs 1/3-1/2 the lemma length;

lemmas (3.2)4.25-5 mm, lanceolate to narrowly lanceolate, distinctly keeled, glabrous, moderately to densely, infrequently sparsely, scabrous, lateral veins prominent, margins narrowly scarious, glabrous, apices acute to narrowly acute;

paleas 3/4 as long as to subequal to the lemmas, keels scabrous, intercostal regions scabrous;

anthers (1.4)1.75-2.6 mm, or vestigial (0.1-0.2 mm).

2n

= unknown.

= unknown.

Poa howellii

Poa diaboli

Distribution
from FNA
CA; OR; WA; BC
[WildflowerSearch map]
[BONAP county map]
[BONAP county map]
Discussion

Poa howellii grows primarily on rocky banks and wooded slopes, from the coastal ranges of southern British Columbia to southern California. It differs from P. bolanderi (see above) in having puberulent, rather than smooth or scabrous, lemmas, and in growing at lower elevations, mostly from near sea level to 1000 m.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Poa diaboli, which is sequentially gynomonoecious, is endemic to upper shaly slopes, in soft coastal scrub and openings in Bishop Pine stands, in the coastal mountains of San Luis Obispo County, California. It is closely related to P. confinis (see previous), from which it differs by a suite of characters. The two species are also ecologically and geographically distinct.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 24, p. 534. FNA vol. 24, p. 552.
Parent taxa Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Homalopoa Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Madropoa > subsect. Madropoa
Sibling taxa
P. abbreviata, P. alpina, P. alsodes, P. ammophila, P. annua, P. arachnifera, P. arctica, P. arida, P. arnowiae, P. atropurpurea, P. autumnalis, P. bigelovii, P. bolanderi, P. bulbosa, P. chaixii, P. chambersii, P. chapmaniana, P. compressa, P. confinis, P. curtifolia, P. cusickii, P. cuspidata, P. diaboli, P. douglasii, P. eminens, P. fendleriana, P. glauca, P. hartzii, P. infirma, P. interior, P. keckii, P. kelloggii, P. laxa, P. laxa × glauca, P. laxiflora, P. leibergii, P. leptocoma, P. lettermanii, P. macrantha, P. macrocalyx, P. marcida, P. napensis, P. nemoralis, P. nervosa, P. occidentalis, P. paludigena, P. palustris, P. paucispicula, P. piperi, P. porsildii, P. pratensis, P. pringlei, P. pseudoabbreviata, P. reflexa, P. rhizomata, P. saltuensis, P. secunda, P. sierrae, P. stebbinsii, P. stenantha, P. strictiramea, P. sublanata, P. suksdorfii, P. supina, P. sylvestris, P. tenerrima, P. tracyi, P. trivialis, P. unilateralis, P. wheeleri, P. wolfii, P. ×gaspensis, P. ×limosa, P. ×nematophylla
P. abbreviata, P. alpina, P. alsodes, P. ammophila, P. annua, P. arachnifera, P. arctica, P. arida, P. arnowiae, P. atropurpurea, P. autumnalis, P. bigelovii, P. bolanderi, P. bulbosa, P. chaixii, P. chambersii, P. chapmaniana, P. compressa, P. confinis, P. curtifolia, P. cusickii, P. cuspidata, P. douglasii, P. eminens, P. fendleriana, P. glauca, P. hartzii, P. howellii, P. infirma, P. interior, P. keckii, P. kelloggii, P. laxa, P. laxa × glauca, P. laxiflora, P. leibergii, P. leptocoma, P. lettermanii, P. macrantha, P. macrocalyx, P. marcida, P. napensis, P. nemoralis, P. nervosa, P. occidentalis, P. paludigena, P. palustris, P. paucispicula, P. piperi, P. porsildii, P. pratensis, P. pringlei, P. pseudoabbreviata, P. reflexa, P. rhizomata, P. saltuensis, P. secunda, P. sierrae, P. stebbinsii, P. stenantha, P. strictiramea, P. sublanata, P. suksdorfii, P. supina, P. sylvestris, P. tenerrima, P. tracyi, P. trivialis, P. unilateralis, P. wheeleri, P. wolfii, P. ×gaspensis, P. ×limosa, P. ×nematophylla
Synonyms P. bolanderi var. howellii
Name authority Vasey & Scribn. Soreng & D.J. Keil
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