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Howell's blue grass

Bernardino bluegrass, San Bernardino blue grass

Habit Plants usually annual, rarely longer-lived; densely tufted, tuft bases narrow; not stoloniferous, not rhizomatous. Plants perennial; loosely tufted, rhizomatous.
Culms

(10) 25-80(120) cm tall, 0.4-1.75 mm thick, usually erect;

nodes terete, usually 1-2 exserted.

10-55 cm, erect or the bases decumbent, terete or weakly compressed;

nodes terete, not exserted.

Sheaths

closed for 1/2 - 7/8 their length, usually weakly compressed and keeled, usually scabrous, rarely smooth;

ligules 1.5-5(10) mm, smooth or scabrous, acute;

blades 1-7(10) mm wide, flat, lax, soft, finely scabrous, apices narrowly prow-shaped, cauline blades 2-10 cm.

closed for about 1/3 their length, terete, smooth, glabrous, bases of basal sheaths glabrous, distal sheath lengths 1.5-7.5 times blade lengths;

collars smooth, glabrous;

ligules 1-2 mm, smooth or sparsely scabrous, apices truncate to obtuse;

innovation blades similar to the cauline blades, adaxial surfaces nearly smooth, glabrous on and between the veins;

cauline blades fairly strongly reduced in length distally, 1-3 mm wide, folded to involute, moderately thick, moderately firm, abaxial surfaces smooth, apices narrowly prow-shaped, flag leaf blades 1-5.5 cm.

Basal branching

intravaginal.

extra- and intravaginal.

Panicles

10-25(30) cm, erect, eventually open, with (1)3-5(7) branches per node;

branches eventually spreading or reflexed, fairly straight, angled, angles usually moderately to densely scabrous, rarely sparsely scabrous.

3-7 cm, erect, lanceoloid to ovoid, congested, with 20-70 spikelets;

nodes with 1-2 branches;

branches 0.5-3 cm, erect, terete, usually smooth, infrequently sparsely scabrous distally, with 3-12 spikelets.

Spikelets

(2)4-6 mm, laterally compressed, with 2-5 florets;

rachilla internodes about 1 mm, smooth, usually softly puberulent, infrequently glabrous.

3.5-5.5 mm, lengths to 3 times widths, laterally compressed, very compact, not sexually dimorphic;

florets 2-5;

rachilla internodes about 0.5 mm, smooth, glabrous.

Glumes

slightly unequal, lanceolate, distinctly keeled, keels and sometimes the lateral veins sparsely to moderately scabrous;

lower glumes 1-3-veined;

upper glumes shorter than or subequal to the lowest lemmas;

calluses of some or all florets sparsely webbed;

lemmas 2.5-3.5 mm, lanceolate to narrowly lanceolate, distinctly keeled, crisply puberulent proximally, hairs evenly distributed, finely scabrous distally, lateral veins obscure to prominent, margins narrowly hyaline, glabrous, apices narrowly acute, infrequently anthocyanic;

palea keels sparsely scabrous, glabrous or softly puberulent at midlength, intercostal regions usually softly puberulent;

anthers 3, 0.2-1 mm.

broadly lanceolate, distinctly shorter than the adjacent lemmas, distinctly keeled, keels smooth or sparsely scabrous;

lower glumes 3-veined;

calluses glabrous;

lemmas 2.5-3.5 mm, lanceolate, usually purplish, distinctly keeled, glabrous, smooth, margins glabrous, apices acute;

palea keels scabrous, intercostal regions glabrous;

anthers vestigial (0.1-0.2 mm) or 1.5-2 mm.

2n

= unknown.

= 28.

Poa howellii

Poa atropurpurea

Distribution
from FNA
CA; OR; WA; BC
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[BONAP county map]
Discussion

Poa howellii grows primarily on rocky banks and wooded slopes, from the coastal ranges of southern British Columbia to southern California. It differs from P. bolanderi (see above) in having puberulent, rather than smooth or scabrous, lemmas, and in growing at lower elevations, mostly from near sea level to 1000 m.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Poa atropurpurea is a rare dioecious endemic of mesic upland meadows in southern California. It is federally listed as endangered.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 24, p. 534. FNA vol. 24, p. 554.
Parent taxa Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Homalopoa Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Madropoa > subsect. Madropoa
Sibling taxa
P. abbreviata, P. alpina, P. alsodes, P. ammophila, P. annua, P. arachnifera, P. arctica, P. arida, P. arnowiae, P. atropurpurea, P. autumnalis, P. bigelovii, P. bolanderi, P. bulbosa, P. chaixii, P. chambersii, P. chapmaniana, P. compressa, P. confinis, P. curtifolia, P. cusickii, P. cuspidata, P. diaboli, P. douglasii, P. eminens, P. fendleriana, P. glauca, P. hartzii, P. infirma, P. interior, P. keckii, P. kelloggii, P. laxa, P. laxa × glauca, P. laxiflora, P. leibergii, P. leptocoma, P. lettermanii, P. macrantha, P. macrocalyx, P. marcida, P. napensis, P. nemoralis, P. nervosa, P. occidentalis, P. paludigena, P. palustris, P. paucispicula, P. piperi, P. porsildii, P. pratensis, P. pringlei, P. pseudoabbreviata, P. reflexa, P. rhizomata, P. saltuensis, P. secunda, P. sierrae, P. stebbinsii, P. stenantha, P. strictiramea, P. sublanata, P. suksdorfii, P. supina, P. sylvestris, P. tenerrima, P. tracyi, P. trivialis, P. unilateralis, P. wheeleri, P. wolfii, P. ×gaspensis, P. ×limosa, P. ×nematophylla
P. abbreviata, P. alpina, P. alsodes, P. ammophila, P. annua, P. arachnifera, P. arctica, P. arida, P. arnowiae, P. autumnalis, P. bigelovii, P. bolanderi, P. bulbosa, P. chaixii, P. chambersii, P. chapmaniana, P. compressa, P. confinis, P. curtifolia, P. cusickii, P. cuspidata, P. diaboli, P. douglasii, P. eminens, P. fendleriana, P. glauca, P. hartzii, P. howellii, P. infirma, P. interior, P. keckii, P. kelloggii, P. laxa, P. laxa × glauca, P. laxiflora, P. leibergii, P. leptocoma, P. lettermanii, P. macrantha, P. macrocalyx, P. marcida, P. napensis, P. nemoralis, P. nervosa, P. occidentalis, P. paludigena, P. palustris, P. paucispicula, P. piperi, P. porsildii, P. pratensis, P. pringlei, P. pseudoabbreviata, P. reflexa, P. rhizomata, P. saltuensis, P. secunda, P. sierrae, P. stebbinsii, P. stenantha, P. strictiramea, P. sublanata, P. suksdorfii, P. supina, P. sylvestris, P. tenerrima, P. tracyi, P. trivialis, P. unilateralis, P. wheeleri, P. wolfii, P. ×gaspensis, P. ×limosa, P. ×nematophylla
Synonyms P. bolanderi var. howellii
Name authority Vasey & Scribn. Scribn.
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