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glaucous blue grass, timberline bluegrass, white bluegrass

Hooker's bluegrass, Wheeler's blue grass

Habit Plants perennial; usually glaucous; densely tufted, not stoloniferous, not rhizomatous. Plants perennial; densely to loosely tufted or with solitary shoots, shortly rhizomatous.
Culms

5-40(80) cm, erect to spreading, straight, wiry, bases straight or slightly decumbent;

nodes terete or slightly compressed, usually 0-1 exserted, top node at 1/10-1/3 the culm length.

35-80 cm, erect or the bases decumbent, terete or weakly compressed;

nodes terete, 1-2 exserted.

Sheaths

closed for 1/10 – 1/5 their length, terete, bases of basal sheaths glabrous or sparsely minutely hairy, hairs 0.1-0.2 mm, distal sheath lengths 1.1-4 times blade lengths;

ligules 1-4(5) mm, sparsely to densely scabrous, apices obtuse to acute, minutely ciliolate;

blades 0.8-2.5 mm wide, flat or folded, thin, soft, appressed or abruptly ascending to spreading, straight, apices narrowly prow-shaped.

closed for 1/3-1/4 their length, terete to slightly compressed, at least some proximal sheaths densely retrorsely scabrous, hispidulous, or softly puberulent for the upper 1/4 of their length, bases of basal sheaths glabrous, distal sheath lengths (1.4)1.7-4.6(6.2) times blade lengths;

collars of proximal leaves glabrous or with hairs the same length as those of their sheaths;

ligules 0.5-2 mm, smooth or scabrous, sometimes puberulent, truncate, those of the lower culm and innovation leaves 0.5-1.5 mm, abaxial surfaces scabrous to softly puberulent, truncate;

innovation blades folded or involute, infrequently flat, moderately thick, soft, adaxial surfaces usually densely scabrous to hispidulous;

cauline blades 2-3.5 mm wide, flat or folded, smooth or sparsely scabrous, glabrous or hispidulous, apices narrowly to broadly prow-shaped, blades gradually reduced distally or the middle blades longest, flag leaf blades 1-10 cm long.

Basal branching

all or mostly extravaginal.

mainly extra-vaginal.

Panicles

1-10(20) cm, lengths 3-5 times widths at maturity, rarely racemelike with branches of irregular length, erect, narrowly lanceoloid to ovoid, contracted to somewhat open, sparse, proximal internodes shorter than 1.5(4) cm;

nodes with 2-3(5) branches;

branches erect, ascending or weakly spreading, fairly straight, short, stout, angled, angles moderately to densely scabrous, rarely only scabrous distally, glaucous;

pedicels usually shorter than the spikelets.

5-12(18) cm, erect or nodding, ovoid to pyramidal, loosely contracted to open, with 20-70 spikelets, proximal internodes usually shorter than 3.5 cm;

nodes with 2-5 branches;

branches (1)1.7-6.5 cm, ascending to spreading or reflexed, lax, terete or weakly angled, sparsely to moderately scabrous, with 2-8(12) spikelets.

Spikelets

3-7(9) mm, lengths 2-3 times widths, laterally compressed, rarely bulbiferous, usually glaucous;

florets 2-5, rarely bulb-forming;

rachilla internodes to 1.2 mm, smooth, muriculate, or scabrous, glabrous or sparsely to densely hispidulous or puberulent.

5.5-10 mm, lengths to 3.5 times widths, laterally compressed, not sexually dimorphic;

florets 2-7;

rachilla internodes smooth or scabrous, glabrous or sparsely to densely hispidulous.

Glumes

subequal, narrowly to broadly lanceolate, distinctly keeled, keels smooth or sparsely scabrous, apices acute;

lower glumes 3-veined;

upper glumes 2-3.8(5.2) mm, lengths usually more than 4.1 times widths, distinctly shorter to subequal to the lowest lemmas;

calluses glabrous or webbed, webs from minute to more than 1/2 the lemma length;

lemmas 2.5-4 mm, lanceolate to broadly lanceolate, distinctly keeled, keels and marginal veins short-villous, lateral veins obscure, usually sparsely softly puberulent to short-villous, intercostal regions smooth, sometimes weakly muriculate, glabrous or puberulent, margins glabrous, apices usually partially bronze-colored, obtuse or acute;

palea keels scabrous, glabrous or softly puberulent at midlength, intercostal regions glabrous or softly puberulent;

anthers (1)1.2-2.5 mm, mature sacs 0.2 mm wide, rarely aborted late in development.

1/4 - 2/3(3/4) as long as the adjacent lemmas, lanceolate, distinctly keeled;

lower glumes 1-3-veined, 1/4-1/2 as long as the adjacent lemmas;

calluses glabrous;

lemmas 3-6 mm, lanceolate, distinctly keeled, keels and marginal veins glabrous or softly puberulent to short-villous, intercostal regions glabrous or hispidulous, infrequently puberulent, smooth or finely scabrous, lateral veins obscure to moderately prominent, margins glabrous, apices acute;

palea keels scabrous, intercostal regions glabrous;

anthers usually vestigial (0.1-0.2 mm) or aborted late in development and up to 2 mm, rarely normal.

2n

= 34, 42, 44, 47, 48, 49, 50, 56, 56, 57, 58, 60, 63, 64, 65, 70, 75, 78, ca. 100.

= 56, 61, 62, 63, 64, 66, 61, 70, ca. 74, 75, 79, 80, 81, 87, 89, 90, 91.

Poa glauca

Poa wheeleri

Distribution
from FNA
AK; AZ; CA; CO; ID; ME; MI; MN; MT; NH; NM; NV; NY; OR; PA; SD; UT; VT; WA; WI; WY; AB; BC; MB; NB; NL; NS; NT; NU; ON; QC; SK; YT; Greenland
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; CO; ID; MT; NM; NV; OR; UT; WA; WY; AB; BC; MB; SK
[WildflowerSearch map]
[BONAP county map]
Discussion

Poa glauca is a common, highly variable, circumboreal, boreal forest to alpine and high arctic species. It grows from Alaska to Greenland, south to California and New Mexico in the west, and through Canada and the northeastern United States in the east. It also grows at scattered locations in Patagonia. It generally favors dry habitats and tolerates disturbance well. It can be distinguished from P. nemoralis (p. 574) and P. interior (see previous) by its longer ligules, lower top culm node, and wider glumes and lemmas. It can be difficult to distinguish from P. laxa subsp. banffiana (p. 570). Poa glauca is often confused in herbaria with P. abbreviata subsp. pattersonii (p. 582). It differs in having primarily extravaginal branching and, usually, longer anthers. It hybridizes with P. laxa, forming P. laxa x glauca (p. 572). It is also known to hybridize with P. hartzii (p. 589), and is suspected to hybridize with P. arctica (p. 529) and P. secunda (p. 586). It is highly polyploid, and presumed to be highly apomictic.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Poa wheeleri is common at mid- to high elevations, generally on the east side of the coastal mountains from British Columbia to California, and from Manitoba to New Mexico. It generally grows in submesic coniferous forests to subalpine habitats. Most plants have densely retrorsely pubescent or scabrous sheaths, involute innovation blades that are pubescent adaxially, and pistillate florets.

Poa wheeleri, a high polyploid apomictic species, probably arose from hybridization between P. cusickii (p. 559) and another member of the Poa nervosa complex. It resembles P. rhizomata (see next) and P. chambersii (p. 548) more than P. nervosa sensu stricto (see previous). It differs from P. chambersii in having at least some proximal sheaths that are densely retrorsely scabrous or pubescent (sometimes obscurely so), and folded or involute innovation blades that are scabrous to hispidulous on the adaxial surfaces. For a comparison with P. nervosa, see p. 545. Natural hybrids have been found between P. wheeleri and P. pratensis (p. 522).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. All or some spikelets bulbiferous
var. pekulnejensis
1. Spikelets not bulbiferous.
→ 2
2. Calluses usually webbed, sometimes glabrous; lemmas glabrous or hairy between the veins
subsp. glauca
2. Calluses glabrous; lemmas hairy between the veins
subsp. rupicola
Source FNA vol. 24, p. 576. FNA vol. 24, p. 546.
Parent taxa Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Stenopoa Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Madropoa > subsect. Poa
Sibling taxa
P. abbreviata, P. alpina, P. alsodes, P. ammophila, P. annua, P. arachnifera, P. arctica, P. arida, P. arnowiae, P. atropurpurea, P. autumnalis, P. bigelovii, P. bolanderi, P. bulbosa, P. chaixii, P. chambersii, P. chapmaniana, P. compressa, P. confinis, P. curtifolia, P. cusickii, P. cuspidata, P. diaboli, P. douglasii, P. eminens, P. fendleriana, P. hartzii, P. howellii, P. infirma, P. interior, P. keckii, P. kelloggii, P. laxa, P. laxa × glauca, P. laxiflora, P. leibergii, P. leptocoma, P. lettermanii, P. macrantha, P. macrocalyx, P. marcida, P. napensis, P. nemoralis, P. nervosa, P. occidentalis, P. paludigena, P. palustris, P. paucispicula, P. piperi, P. porsildii, P. pratensis, P. pringlei, P. pseudoabbreviata, P. reflexa, P. rhizomata, P. saltuensis, P. secunda, P. sierrae, P. stebbinsii, P. stenantha, P. strictiramea, P. sublanata, P. suksdorfii, P. supina, P. sylvestris, P. tenerrima, P. tracyi, P. trivialis, P. unilateralis, P. wheeleri, P. wolfii, P. ×gaspensis, P. ×limosa, P. ×nematophylla
P. abbreviata, P. alpina, P. alsodes, P. ammophila, P. annua, P. arachnifera, P. arctica, P. arida, P. arnowiae, P. atropurpurea, P. autumnalis, P. bigelovii, P. bolanderi, P. bulbosa, P. chaixii, P. chambersii, P. chapmaniana, P. compressa, P. confinis, P. curtifolia, P. cusickii, P. cuspidata, P. diaboli, P. douglasii, P. eminens, P. fendleriana, P. glauca, P. hartzii, P. howellii, P. infirma, P. interior, P. keckii, P. kelloggii, P. laxa, P. laxa × glauca, P. laxiflora, P. leibergii, P. leptocoma, P. lettermanii, P. macrantha, P. macrocalyx, P. marcida, P. napensis, P. nemoralis, P. nervosa, P. occidentalis, P. paludigena, P. palustris, P. paucispicula, P. piperi, P. porsildii, P. pratensis, P. pringlei, P. pseudoabbreviata, P. reflexa, P. rhizomata, P. saltuensis, P. secunda, P. sierrae, P. stebbinsii, P. stenantha, P. strictiramea, P. sublanata, P. suksdorfii, P. supina, P. sylvestris, P. tenerrima, P. tracyi, P. trivialis, P. unilateralis, P. wolfii, P. ×gaspensis, P. ×limosa, P. ×nematophylla
Subordinate taxa
P. glauca subsp. glauca, P. glauca subsp. rupicola, P. glauca var. pekulnejensis
Synonyms P. nervosa var. wheeleri, P. curta
Name authority Vahl Vasey
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