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glaucous blue grass, timberline bluegrass, white bluegrass

Cusick's bluegrass

Habit Plants perennial; usually glaucous; densely tufted, not stoloniferous, not rhizomatous. Plants perennial; usually densely tufted, rarely moderately densely tufted, usually neither rhizomatous nor stoloniferous, infrequently short-rhizomatous or stoloniferous, rarely with distinct rhizomes.
Culms

5-40(80) cm, erect to spreading, straight, wiry, bases straight or slightly decumbent;

nodes terete or slightly compressed, usually 0-1 exserted, top node at 1/10-1/3 the culm length.

10-60(70) cm tall, 0.5-1.8 mm thick, erect or the bases decumbent, terete or weakly compressed;

nodes terete, 0-2 exserted.

Sheaths

closed for 1/10 – 1/5 their length, terete, bases of basal sheaths glabrous or sparsely minutely hairy, hairs 0.1-0.2 mm, distal sheath lengths 1.1-4 times blade lengths;

ligules 1-4(5) mm, sparsely to densely scabrous, apices obtuse to acute, minutely ciliolate;

blades 0.8-2.5 mm wide, flat or folded, thin, soft, appressed or abruptly ascending to spreading, straight, apices narrowly prow-shaped.

closed for 1/4-3/4 their length, terete, smooth or scabrous, glabrous, bases of basal sheaths glabrous, distal sheath lengths 1.6-10 times blade lengths;

collars smooth or scabrous, glabrous;

ligules of cauline leaves 1-3(6) mm, smooth or scabrous, truncate to acute, ligules of the innovation leaves 0.2-0.5(2.5) mm, scabrous, usually truncate;

innovation blades sometimes distinctly different from the cauline blades, 0.5-2 mm wide, involute, moderately thick, moderately firm, adaxial surfaces usually densely scabrous or hispidulous to softly puberulent, infrequently nearly smooth and glabrous;

cauline blades subequal or the midcauline blades longest or the blades gradually reduced in length distally, 0.5-3 mm wide, flat, folded, or involute, usually thin, usually withering, abaxial surfaces smooth or scabrous, apices narrowly to broadly prow-shaped, flag leaf blades 0.5-5(6) cm.

Basal branching

all or mostly extravaginal.

intravaginal or intra- and extravaginal.

Panicles

1-10(20) cm, lengths 3-5 times widths at maturity, rarely racemelike with branches of irregular length, erect, narrowly lanceoloid to ovoid, contracted to somewhat open, sparse, proximal internodes shorter than 1.5(4) cm;

nodes with 2-3(5) branches;

branches erect, ascending or weakly spreading, fairly straight, short, stout, angled, angles moderately to densely scabrous, rarely only scabrous distally, glaucous;

pedicels usually shorter than the spikelets.

2-10(12) cm, usually erect, contracted or loosely contracted, narrowly lanceoloid to ovoid, congested or moderately congested, with 10-100 spikelets and 1-3(5) branches per node;

branches 0.5-4(5) cm, erect or steeply ascending, fairly straight, slender to stout, terete to angled, smooth or scabrous, with 1-15 spikelets.

Spikelets

3-7(9) mm, lengths 2-3 times widths, laterally compressed, rarely bulbiferous, usually glaucous;

florets 2-5, rarely bulb-forming;

rachilla internodes to 1.2 mm, smooth, muriculate, or scabrous, glabrous or sparsely to densely hispidulous or puberulent.

(3)4-10 mm, lengths to 3 times widths, broadly lanceolate to narrowly ovate, laterally compressed, not sexually dimorphic;

florets 2-6;

rachilla internodes 0.5-1.2 mm, smooth or scabrous.

Glumes

subequal, narrowly to broadly lanceolate, distinctly keeled, keels smooth or sparsely scabrous, apices acute;

lower glumes 3-veined;

upper glumes 2-3.8(5.2) mm, lengths usually more than 4.1 times widths, distinctly shorter to subequal to the lowest lemmas;

calluses glabrous or webbed, webs from minute to more than 1/2 the lemma length;

lemmas 2.5-4 mm, lanceolate to broadly lanceolate, distinctly keeled, keels and marginal veins short-villous, lateral veins obscure, usually sparsely softly puberulent to short-villous, intercostal regions smooth, sometimes weakly muriculate, glabrous or puberulent, margins glabrous, apices usually partially bronze-colored, obtuse or acute;

palea keels scabrous, glabrous or softly puberulent at midlength, intercostal regions glabrous or softly puberulent;

anthers (1)1.2-2.5 mm, mature sacs 0.2 mm wide, rarely aborted late in development.

lanceolate, distinctly keeled;

lower glumes 3-veined, distinctly shorter than the lowest lemmas;

calluses glabrous or diffusely webbed, hairs less than 1/4 the lemma length;

lemmas (3)4-7 mm, lanceolate to broadly lanceolate, distinctly keeled, membranous to thinly membranous, smooth or sparsely to densely scabrous, glabrous or the keels and/or marginal veins puberulent proximally, lateral veins obscure to prominent, margins glabrous, apices acute;

palea keels scabrous, intercostal regions glabrous;

anthers vestigial (0.1-0.2 mm), aborted late in development, or 2-3.5 mm.

2n

= 34, 42, 44, 47, 48, 49, 50, 56, 56, 57, 58, 60, 63, 64, 65, 70, 75, 78, ca. 100.

= 28, 28+11, 56, 56+11, 59, ca. 70.

Poa glauca

Poa cusickii

Distribution
from FNA
AK; AZ; CA; CO; ID; ME; MI; MN; MT; NH; NM; NV; NY; OR; PA; SD; UT; VT; WA; WI; WY; AB; BC; MB; NB; NL; NS; NT; NU; ON; QC; SK; YT; Greenland
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; CO; ID; MT; ND; NV; OR; UT; WA; WY; AB; BC; MB; SK; YT
[WildflowerSearch map]
[BONAP county map]
Discussion

Poa glauca is a common, highly variable, circumboreal, boreal forest to alpine and high arctic species. It grows from Alaska to Greenland, south to California and New Mexico in the west, and through Canada and the northeastern United States in the east. It also grows at scattered locations in Patagonia. It generally favors dry habitats and tolerates disturbance well. It can be distinguished from P. nemoralis (p. 574) and P. interior (see previous) by its longer ligules, lower top culm node, and wider glumes and lemmas. It can be difficult to distinguish from P. laxa subsp. banffiana (p. 570). Poa glauca is often confused in herbaria with P. abbreviata subsp. pattersonii (p. 582). It differs in having primarily extravaginal branching and, usually, longer anthers. It hybridizes with P. laxa, forming P. laxa x glauca (p. 572). It is also known to hybridize with P. hartzii (p. 589), and is suspected to hybridize with P. arctica (p. 529) and P. secunda (p. 586). It is highly polyploid, and presumed to be highly apomictic.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Poa cusickii grows in rich meadows in sagebrush scrub to rocky alpine slopes, from the southwestern Yukon Territory to Manitoba and North Dakota, south to central California and eastern Colorado. It is gynodioecious or dioecious.

Sexually reproducing plants of Poa cusickii subspp. cusickii and pallida grow in different geographic areas, but pistillate plants of these two subspecies have overlapping ranges. Only pistillate plants are known in Poa cusickii subspp. epilis and purpurascens. All the alpine plants studied were pistillate.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. All or some spikelets bulbiferous
var. pekulnejensis
1. Spikelets not bulbiferous.
→ 2
2. Calluses usually webbed, sometimes glabrous; lemmas glabrous or hairy between the veins
subsp. glauca
2. Calluses glabrous; lemmas hairy between the veins
subsp. rupicola
1. Panicle branches smooth or slightly scabrous, or the basal blades more than 1.5 mm wide and flat or folded; cauline blades more than 1.5 mm wide, often flat; some basal branching extravaginal; lemmas and calluses sometimes sparsely puberulent.
→ 2
2. Lemmas usually glabrous, rarely plants from the Rocky Mountains with puberulent keels and marginal veins; calluses glabrous; panicles erect, usually contracted; branches smooth to slightly scabrous
subsp. epilis
2. Lemmas rarely completely glabrous, at least some florets v/ith sparsely puberulent keels, the marginal veins glabrous or puberulent; calluses frequendy with a sparse, short web; panicles somewhat lax and loosely contracted; branches smooth or sparsely to moderately scabrous
subsp. purpurascens
1. Panicle branches moderately to strongly scabrous; basal and cauline blades usually less than 1.5 mm wide, involute, rarely flat or folded; basal branching intravaginal; lemmas and calluses glabrous.
→ 3
3. Panicle branches longer than 1.7 cm in at least some panicles; panicles open or contracted
subsp. cusickii
3. Panicle branches up to 1.7 cm long, stout; panicles contracted
subsp. pallida
Source FNA vol. 24, p. 576. FNA vol. 24, p. 559.
Parent taxa Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Stenopoa Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Madropoa > subsect. Epiles
Sibling taxa
P. abbreviata, P. alpina, P. alsodes, P. ammophila, P. annua, P. arachnifera, P. arctica, P. arida, P. arnowiae, P. atropurpurea, P. autumnalis, P. bigelovii, P. bolanderi, P. bulbosa, P. chaixii, P. chambersii, P. chapmaniana, P. compressa, P. confinis, P. curtifolia, P. cusickii, P. cuspidata, P. diaboli, P. douglasii, P. eminens, P. fendleriana, P. hartzii, P. howellii, P. infirma, P. interior, P. keckii, P. kelloggii, P. laxa, P. laxa × glauca, P. laxiflora, P. leibergii, P. leptocoma, P. lettermanii, P. macrantha, P. macrocalyx, P. marcida, P. napensis, P. nemoralis, P. nervosa, P. occidentalis, P. paludigena, P. palustris, P. paucispicula, P. piperi, P. porsildii, P. pratensis, P. pringlei, P. pseudoabbreviata, P. reflexa, P. rhizomata, P. saltuensis, P. secunda, P. sierrae, P. stebbinsii, P. stenantha, P. strictiramea, P. sublanata, P. suksdorfii, P. supina, P. sylvestris, P. tenerrima, P. tracyi, P. trivialis, P. unilateralis, P. wheeleri, P. wolfii, P. ×gaspensis, P. ×limosa, P. ×nematophylla
P. abbreviata, P. alpina, P. alsodes, P. ammophila, P. annua, P. arachnifera, P. arctica, P. arida, P. arnowiae, P. atropurpurea, P. autumnalis, P. bigelovii, P. bolanderi, P. bulbosa, P. chaixii, P. chambersii, P. chapmaniana, P. compressa, P. confinis, P. curtifolia, P. cuspidata, P. diaboli, P. douglasii, P. eminens, P. fendleriana, P. glauca, P. hartzii, P. howellii, P. infirma, P. interior, P. keckii, P. kelloggii, P. laxa, P. laxa × glauca, P. laxiflora, P. leibergii, P. leptocoma, P. lettermanii, P. macrantha, P. macrocalyx, P. marcida, P. napensis, P. nemoralis, P. nervosa, P. occidentalis, P. paludigena, P. palustris, P. paucispicula, P. piperi, P. porsildii, P. pratensis, P. pringlei, P. pseudoabbreviata, P. reflexa, P. rhizomata, P. saltuensis, P. secunda, P. sierrae, P. stebbinsii, P. stenantha, P. strictiramea, P. sublanata, P. suksdorfii, P. supina, P. sylvestris, P. tenerrima, P. tracyi, P. trivialis, P. unilateralis, P. wheeleri, P. wolfii, P. ×gaspensis, P. ×limosa, P. ×nematophylla
Subordinate taxa
P. glauca subsp. glauca, P. glauca subsp. rupicola, P. glauca var. pekulnejensis
P. cusickii subsp. cusickii, P. cusickii subsp. epilis, P. cusickii subsp. pallida, P. cusickii subsp. purpurascens
Name authority Vahl Vasey
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