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Fendler's bluegrass, long-tongue mutton grass, mutton grass, Vasey's mutton grass

Hartz's bluegrass

Habit Plants perennial; densely to loosely tufted, rhizomatous, often weakly so, rhizomes usually short and inconspicuous. Plants perennial; not glaucous; densely to loosely tufted, not rhizomatous,occasionally weakly stoloniferous.
Culms

15-70 cm, sometimes stout, erect or the bases decumbent, terete or weakly compressed;

nodes terete, 0-1 exserted.

10-33(45) cm, usually decumbent, terete;

nodes terete, 0(1) exserted.

Sheaths

closed for about 1/3 their length, terete, smooth or scabrous, glabrous or occasionally retrorsely pubescent, bases of basal sheaths glabrous, distal sheath lengths usually (5)9+ times blade lengths;

collars smooth or scabrous, glabrous or hispidulous;

ligules 0.2-18 mm, smooth or scabrous, decurrent or not, apices truncate to acuminate, ciliolate or glabrous;

innovation blades usually moderately to densely scabrous or hispidulous on and between the veins, infrequently nearly smooth and glabrous;

cauline blades strongly reduced in length distally, (0.5)1-3(4) mm wide, usually involute, moderately thick and firm, infrequently moderately thin, abaxial surfaces usually smooth, infrequently scabrous, apices narrowly prow-shaped, steeply reduced in length distally along the culm, flag leaf blades often absent or very reduced, sometimes to 1(3) cm.

closed for 1/7 – 1/5(1/3) their length, terete, usually lustrous, bases of basal sheaths glabrous;

ligules (1.5)2-7 mm, smooth or sparsely scabrous, margins usually decurrent, apices obtuse to acuminate;

innovation blades similar in texture and shape to those of the culms;

cauline blades 2-9 cm, gradually increasing or decreasing in length upwards, 1.5-3 mm wide, folded to involute, moderately thick, soft, abaxial surfaces smooth, adaxial surfaces smooth or somewhat scabrous, usually glabrous, infrequently sparsely hispidulous, apices narrowly prow-shaped.

Basal branching

mainly intravaginal, usually some extravaginal.

extra- and intravaginal.

Panicles

2-12(30) cm, erect, contracted, narrowly lanceoloid to ovoid, congested, frequently with 100+ spikelets;

nodes with 1-2 branches;

branches 1-8 cm, erect, terete to weakly angled, smooth or scabrous, with 3-15(25) spikelets.

2.5-6(12) cm, erect, narrowly lanceolate, contracted or narrowly ovate in some bulbiferous plants, moderately congested, with 7-40 spikelets;

nodes with (1)2(4) branches;

branches 1-3 cm, erect to ascending, straight, sulcate, smooth or sparsely to moderately scabrous, with 1-10 spikelets in the distal 1/3 – 2/3.

Spikelets

(3)4-8(12) mm, lengths to 3 times widths, broadly lanceolate to ovate, laterally compressed, not sexually dimorphic;

florets 2-7(13);

rachilla internodes 0.8-1.3 mm, smooth, glabrous or hairy, hairs to 0.3 mm.

4.8-7.4 mm, lengths 3.5-4 times widths, lanceolate, weakly laterally compressed, sometimes bulbiferous, lustrous;

florets (2)3-5(6), normal or bulb-forming;

rachilla internodes 0.8-2 mm, smooth, sometimes sparsely hispidulous.

Glumes

lanceolate, distinctly keeled;

lower glumes 1-3-veined, distinctly shorter than the lowest lemmas;

calluses glabrous;

lemmas 3-6 mm, lanceolate, distinctly keeled, keels, marginal veins, and lateral veins glabrous or short- to long-villous or softly puberulent, lateral veins moderately prominent, intercostal regions softly puberulent or glabrous, smooth or sparsely scabrous, margins glabrous, apices acute;

palea keels scabrous, sometimes softly puberulent or long-villous at midlength, hairs to 0.4+ mm;

anthers vestigial (0.1-0.2 mm) or 2-3 mm.

mostly broadly scarious, somewhat lustrous, keels indistinct, smooth or sparsely scabrous distally;

lower glumes 3-veined;

upper glumes frequently exceeding the lowest lemmas;

calluses glabrous or with a crown of hairs, hairs to 2 mm;

lemmas (3.3)3.5-7 mm, lanceolate, usually weakly keeled, more or less evenly and somewhat loosely to densely hairy over the proximal 1/3-1/2, hairs usually longer than 0.5 mm, sparsely scabrous in the middle 1/3, smooth distally, lateral veins obscure, margins weakly inrolled, broadly scarious, glabrous, apices long-scarious, acute to shortly obtuse, often erose, often bronze-colored below the apices;

palea keels sparsely scabrous, softly puberulent at midlength, intercostal regions softly puberulent;

anthers usually all aborted late in development and 0.8-1.8 mm, infrequently well developed and 2-2.8 mm.

2n

= 28+11, 56, 56-58, 58-64.

= 63, 70.

Poa fendleriana

Poa hartzii

Distribution
from FNA
AZ; CA; CO; ID; MT; ND; NE; NM; NV; OK; OR; SD; TX; UT; WA; WY; AB; BC; MB
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; NT; NU; QC; Greenland
[BONAP county map]
Discussion

Poa fendleriana grows on rocky to rich slopes in sagebrush-scrub, interior chaparral, and southern (rarely northern) high plains grasslands to forests, and from desert hills to low alpine habitats. Its range extends from British Columbia to Manitoba and south to Mexico. It is one of the best spring fodder grasses in the eastern Great Basin, Colorado plateaus, and southern Rocky Mountains. It is dioecious. Each of the subspecies has regions of sexual reproduction in which staminate plants are common within populations, and extensive regions where only apomictic, pistillate plants are found. The sexual populations set little seed; the apomictic populations are highly fecund.

Poa fendleriana hybridizes with Poa cusickii subsp. pallida (p. 560). The hybrids are called P. xnematophylla (p. 562).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Poa hartzii grows only in the high arctic. It generally grows on open ground, on sandy or clayey soils, or on slumping slopes of old marine terraces. It carries two chloroplast genomes within its populations; one of these links it to P. secunda (p. 586) and P. ammophila (see next), the other to P. glauca (p. 576). Morphologically, it is closest to P. secunda and P. ammophila.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Lemma keels and marginal veins glabrous or almost so
subsp. albescens
1. Lemma keels and marginal veins conspicuously hairy.
→ 2
2. Ligules of the middle cauline leaves 0.2-1.2 (1.5) mm long, not decurrent, usually scabrous, apices truncate to rounded, upper margins ciliolate or scabrous
subsp. fendleriana
2. Ligules of the middle cauline leaves (1.5)1.8-18 mm long, decurrent, usually smooth to sparsely scabrous, apices obtuse to acuminate, upper margins usually smooth, glabrous
subsp. longiligula
1. Spikelets bulbiferous
subsp. vrangelica
1. Spikelets not bulbiferous.
→ 2
2. Lemmas 5.5-7 mm long; anthers well developed, 2-2.8 mm long
subsp. alaskana
2. Lemmas 3.3-5.4 mm long; anthers usually aborted and shorter than 1.5 mm
subsp. hartzii
Source FNA vol. 24, p. 556. FNA vol. 24, p. 589.
Parent taxa Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Madropoa > subsect. Madropoa Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Secundae > subsect. Secundae
Sibling taxa
P. abbreviata, P. alpina, P. alsodes, P. ammophila, P. annua, P. arachnifera, P. arctica, P. arida, P. arnowiae, P. atropurpurea, P. autumnalis, P. bigelovii, P. bolanderi, P. bulbosa, P. chaixii, P. chambersii, P. chapmaniana, P. compressa, P. confinis, P. curtifolia, P. cusickii, P. cuspidata, P. diaboli, P. douglasii, P. eminens, P. glauca, P. hartzii, P. howellii, P. infirma, P. interior, P. keckii, P. kelloggii, P. laxa, P. laxa × glauca, P. laxiflora, P. leibergii, P. leptocoma, P. lettermanii, P. macrantha, P. macrocalyx, P. marcida, P. napensis, P. nemoralis, P. nervosa, P. occidentalis, P. paludigena, P. palustris, P. paucispicula, P. piperi, P. porsildii, P. pratensis, P. pringlei, P. pseudoabbreviata, P. reflexa, P. rhizomata, P. saltuensis, P. secunda, P. sierrae, P. stebbinsii, P. stenantha, P. strictiramea, P. sublanata, P. suksdorfii, P. supina, P. sylvestris, P. tenerrima, P. tracyi, P. trivialis, P. unilateralis, P. wheeleri, P. wolfii, P. ×gaspensis, P. ×limosa, P. ×nematophylla
P. abbreviata, P. alpina, P. alsodes, P. ammophila, P. annua, P. arachnifera, P. arctica, P. arida, P. arnowiae, P. atropurpurea, P. autumnalis, P. bigelovii, P. bolanderi, P. bulbosa, P. chaixii, P. chambersii, P. chapmaniana, P. compressa, P. confinis, P. curtifolia, P. cusickii, P. cuspidata, P. diaboli, P. douglasii, P. eminens, P. fendleriana, P. glauca, P. howellii, P. infirma, P. interior, P. keckii, P. kelloggii, P. laxa, P. laxa × glauca, P. laxiflora, P. leibergii, P. leptocoma, P. lettermanii, P. macrantha, P. macrocalyx, P. marcida, P. napensis, P. nemoralis, P. nervosa, P. occidentalis, P. paludigena, P. palustris, P. paucispicula, P. piperi, P. porsildii, P. pratensis, P. pringlei, P. pseudoabbreviata, P. reflexa, P. rhizomata, P. saltuensis, P. secunda, P. sierrae, P. stebbinsii, P. stenantha, P. strictiramea, P. sublanata, P. suksdorfii, P. supina, P. sylvestris, P. tenerrima, P. tracyi, P. trivialis, P. unilateralis, P. wheeleri, P. wolfii, P. ×gaspensis, P. ×limosa, P. ×nematophylla
Subordinate taxa
P. fendleriana subsp. albescens, P. fendleriana subsp. fendleriana, P. fendleriana subsp. longiligula
P. hartzii subsp. alaskana, P. hartzii subsp. hartzii, P. hartzii subsp. vrangelica
Name authority (Steud.) Vasey Gand.
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