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Douglas' bluegrass, sand dune blue grass

Hooker's bluegrass, Wheeler's blue grass

Habit Plants perennial; loosely tufted, rhizomatous and stoloniferous, rhizomes and stolons to 1 m. Plants perennial; densely to loosely tufted or with solitary shoots, shortly rhizomatous.
Culms

(5)10-30 cm tall, 1.2-1.5 mm thick, bases decumbent, terete or weakly compressed, hispidulous beneath the panicles;

nodes terete, 0(1) exserted.

35-80 cm, erect or the bases decumbent, terete or weakly compressed;

nodes terete, 1-2 exserted.

Sheaths

closed for about 1/2 their length, terete, smooth or sparsely to moderately retrorsely scabrous near the collars, glabrous, bases of basal sheaths glabrous, distal sheath lengths 0.9-3.5 times blade lengths;

collars sparsely to moderately retrorsely scabrous, glabrous;

ligules 1-2 mm, scabrous, truncate to obtuse, ciliolate;

innovation blades to 30 cm long, adaxial surfaces moderately to densely scabrous or hispidulous on and between the veins;

cauline blades subequal in length, 1-2 mm wide, involute, moderately thick, moderately firm, arcuate, abaxial surfaces smooth or sparsely scabrous, adaxial surfaces moderately to densely scabrous or hispidulous on and between the veins, apices narrowly prow-shaped, flag leaf blades 1-9 cm.

closed for 1/3-1/4 their length, terete to slightly compressed, at least some proximal sheaths densely retrorsely scabrous, hispidulous, or softly puberulent for the upper 1/4 of their length, bases of basal sheaths glabrous, distal sheath lengths (1.4)1.7-4.6(6.2) times blade lengths;

collars of proximal leaves glabrous or with hairs the same length as those of their sheaths;

ligules 0.5-2 mm, smooth or scabrous, sometimes puberulent, truncate, those of the lower culm and innovation leaves 0.5-1.5 mm, abaxial surfaces scabrous to softly puberulent, truncate;

innovation blades folded or involute, infrequently flat, moderately thick, soft, adaxial surfaces usually densely scabrous to hispidulous;

cauline blades 2-3.5 mm wide, flat or folded, smooth or sparsely scabrous, glabrous or hispidulous, apices narrowly to broadly prow-shaped, blades gradually reduced distally or the middle blades longest, flag leaf blades 1-10 cm long.

Basal branching

mainly intravaginal, some extravaginal.

mainly extra-vaginal.

Panicles

1.5-6 cm, erect, compact, ovoid, contracted, infrequently interrupted, congested, with 15-50 spikelets;

nodes with 1-2 branches, internodes densely hispidulous;

branches 0.5-2 cm, erect, stiff, terete to weakly angled, densely hispidulous, with 1-5 spikelets.

5-12(18) cm, erect or nodding, ovoid to pyramidal, loosely contracted to open, with 20-70 spikelets, proximal internodes usually shorter than 3.5 cm;

nodes with 2-5 branches;

branches (1)1.7-6.5 cm, ascending to spreading or reflexed, lax, terete or weakly angled, sparsely to moderately scabrous, with 2-8(12) spikelets.

Spikelets

7-12 mm, lengths to 3 times widths, laterally compressed, not sexually dimorphic;

florets 3-6;

rachilla internodes usually shorter than 0.5 mm, smooth, glabrous.

5.5-10 mm, lengths to 3.5 times widths, laterally compressed, not sexually dimorphic;

florets 2-7;

rachilla internodes smooth or scabrous, glabrous or sparsely to densely hispidulous.

Glumes

broadly lanceolate, 1/2 as long as to subequal to the adjacent lemmas, distinctly keeled;

lower glumes 3-veined;

upper glumes 4-4.5(7+) mm, 3-veined;

calluses usually with a crown of hairs, sometimes glabrous or diffusely webbed;

lemmas 5-7.5 mm, lanceolate, 5-veined, distinctly keeled, keels, marginal veins, and sometimes the lateral veins short- to long-villous or softly puberulent, rarely glabrous, lateral veins moderately prominent, intercostal regions smooth, glabrous, margins glabrous, apices acute;

palea keels scabrous to pectinate-ciliate, intercostal regions glabrous;

anthers vestigial (0.1-0.2 mm) or (2)2.5-3.5 (4) mm.

1/4 - 2/3(3/4) as long as the adjacent lemmas, lanceolate, distinctly keeled;

lower glumes 1-3-veined, 1/4-1/2 as long as the adjacent lemmas;

calluses glabrous;

lemmas 3-6 mm, lanceolate, distinctly keeled, keels and marginal veins glabrous or softly puberulent to short-villous, intercostal regions glabrous or hispidulous, infrequently puberulent, smooth or finely scabrous, lateral veins obscure to moderately prominent, margins glabrous, apices acute;

palea keels scabrous, intercostal regions glabrous;

anthers usually vestigial (0.1-0.2 mm) or aborted late in development and up to 2 mm, rarely normal.

2n

= 28.

= 56, 61, 62, 63, 64, 66, 61, 70, ca. 74, 75, 79, 80, 81, 87, 89, 90, 91.

Poa douglasii

Poa wheeleri

Distribution
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; CO; ID; MT; NM; NV; OR; UT; WA; WY; AB; BC; MB; SK
[WildflowerSearch map]
[BONAP county map]
Discussion

Poa douglasii is a dioecious endemic that grows on coastal sand dunes in California, a habitat that is being invaded by exotic species. It is rare north of Mendocino. Its hairy rachises distinguish P. douglasii from all other species of Poa in the Flora region. It differs from P. macrantha (see next), which occupies similar habitats, in this and in its usually longer glumes and lemmas.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Poa wheeleri is common at mid- to high elevations, generally on the east side of the coastal mountains from British Columbia to California, and from Manitoba to New Mexico. It generally grows in submesic coniferous forests to subalpine habitats. Most plants have densely retrorsely pubescent or scabrous sheaths, involute innovation blades that are pubescent adaxially, and pistillate florets.

Poa wheeleri, a high polyploid apomictic species, probably arose from hybridization between P. cusickii (p. 559) and another member of the Poa nervosa complex. It resembles P. rhizomata (see next) and P. chambersii (p. 548) more than P. nervosa sensu stricto (see previous). It differs from P. chambersii in having at least some proximal sheaths that are densely retrorsely scabrous or pubescent (sometimes obscurely so), and folded or involute innovation blades that are scabrous to hispidulous on the adaxial surfaces. For a comparison with P. nervosa, see p. 545. Natural hybrids have been found between P. wheeleri and P. pratensis (p. 522).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 24, p. 551. FNA vol. 24, p. 546.
Parent taxa Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Madropoa > subsect. Madropoa Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Madropoa > subsect. Poa
Sibling taxa
P. abbreviata, P. alpina, P. alsodes, P. ammophila, P. annua, P. arachnifera, P. arctica, P. arida, P. arnowiae, P. atropurpurea, P. autumnalis, P. bigelovii, P. bolanderi, P. bulbosa, P. chaixii, P. chambersii, P. chapmaniana, P. compressa, P. confinis, P. curtifolia, P. cusickii, P. cuspidata, P. diaboli, P. eminens, P. fendleriana, P. glauca, P. hartzii, P. howellii, P. infirma, P. interior, P. keckii, P. kelloggii, P. laxa, P. laxa × glauca, P. laxiflora, P. leibergii, P. leptocoma, P. lettermanii, P. macrantha, P. macrocalyx, P. marcida, P. napensis, P. nemoralis, P. nervosa, P. occidentalis, P. paludigena, P. palustris, P. paucispicula, P. piperi, P. porsildii, P. pratensis, P. pringlei, P. pseudoabbreviata, P. reflexa, P. rhizomata, P. saltuensis, P. secunda, P. sierrae, P. stebbinsii, P. stenantha, P. strictiramea, P. sublanata, P. suksdorfii, P. supina, P. sylvestris, P. tenerrima, P. tracyi, P. trivialis, P. unilateralis, P. wheeleri, P. wolfii, P. ×gaspensis, P. ×limosa, P. ×nematophylla
P. abbreviata, P. alpina, P. alsodes, P. ammophila, P. annua, P. arachnifera, P. arctica, P. arida, P. arnowiae, P. atropurpurea, P. autumnalis, P. bigelovii, P. bolanderi, P. bulbosa, P. chaixii, P. chambersii, P. chapmaniana, P. compressa, P. confinis, P. curtifolia, P. cusickii, P. cuspidata, P. diaboli, P. douglasii, P. eminens, P. fendleriana, P. glauca, P. hartzii, P. howellii, P. infirma, P. interior, P. keckii, P. kelloggii, P. laxa, P. laxa × glauca, P. laxiflora, P. leibergii, P. leptocoma, P. lettermanii, P. macrantha, P. macrocalyx, P. marcida, P. napensis, P. nemoralis, P. nervosa, P. occidentalis, P. paludigena, P. palustris, P. paucispicula, P. piperi, P. porsildii, P. pratensis, P. pringlei, P. pseudoabbreviata, P. reflexa, P. rhizomata, P. saltuensis, P. secunda, P. sierrae, P. stebbinsii, P. stenantha, P. strictiramea, P. sublanata, P. suksdorfii, P. supina, P. sylvestris, P. tenerrima, P. tracyi, P. trivialis, P. unilateralis, P. wolfii, P. ×gaspensis, P. ×limosa, P. ×nematophylla
Synonyms P. nervosa var. wheeleri, P. curta
Name authority Nees Vasey
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