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Douglas' bluegrass, sand dune blue grass

delicate blue grass

Habit Plants perennial; loosely tufted, rhizomatous and stoloniferous, rhizomes and stolons to 1 m. Plants perennial; densely tufted, basal leaf tufts 2-8(13) cm, small, narrowly based, not stoloniferous, not rhizomatous.
Culms

(5)10-30 cm tall, 1.2-1.5 mm thick, bases decumbent, terete or weakly compressed, hispidulous beneath the panicles;

nodes terete, 0(1) exserted.

15-50 cm tall, slender, 0.8-0.9 mm thick;

nodes terete, 0-1 exserted.

Sheaths

closed for about 1/2 their length, terete, smooth or sparsely to moderately retrorsely scabrous near the collars, glabrous, bases of basal sheaths glabrous, distal sheath lengths 0.9-3.5 times blade lengths;

collars sparsely to moderately retrorsely scabrous, glabrous;

ligules 1-2 mm, scabrous, truncate to obtuse, ciliolate;

innovation blades to 30 cm long, adaxial surfaces moderately to densely scabrous or hispidulous on and between the veins;

cauline blades subequal in length, 1-2 mm wide, involute, moderately thick, moderately firm, arcuate, abaxial surfaces smooth or sparsely scabrous, adaxial surfaces moderately to densely scabrous or hispidulous on and between the veins, apices narrowly prow-shaped, flag leaf blades 1-9 cm.

closed for 76-74 their length, terete, scabrous, glabrous, bases of basal sheaths glabrous, distal sheath lengths 2.3-7.7 times blade lengths;

collars sparsely scabrous, glabrous;

ligules 0.5-1.5(2.5) mm, scabrous, apices usually truncate to obtuse, sometimes acute, ligules of innovations to 0.5 mm;

innovation blades filiform;

cauline blades gradually reduced in length upwards, 0.4-1.5 mm wide, mostly folded, thin, soft, soon withering, scabrous, apices narrowly prow-shaped.

Basal branching

mainly intravaginal, some extravaginal.

intravaginal.

Panicles

1.5-6 cm, erect, compact, ovoid, contracted, infrequently interrupted, congested, with 15-50 spikelets;

nodes with 1-2 branches, internodes densely hispidulous;

branches 0.5-2 cm, erect, stiff, terete to weakly angled, densely hispidulous, with 1-5 spikelets.

5-15 cm, 1.3-2.2 times the branch lengths, erect, broadly rhomboidal to pyramidal, open, purple, sparse, proximal internodes 1.5-4.2 cm;

nodes with 1-2(5) branches;

branches 3-8.5 cm, widely spreading, capillary, straight, terete to weakly angled, moderately to mostly densely scabrous, with 3-9 spikelets in the distal 73.

Spikelets

7-12 mm, lengths to 3 times widths, laterally compressed, not sexually dimorphic;

florets 3-6;

rachilla internodes usually shorter than 0.5 mm, smooth, glabrous.

5-8 mm, lengths (3.8)4-5 times widths, usually narrowly lanceolate to lanceolate, subterete to weakly laterally compressed, drab, usually strongly anthocyanic;

florets 3-5;

rachilla internodes 1-1.5+ mm, terete or slightly dorsally compressed, muriculate or scabrous.

Glumes

broadly lanceolate, 1/2 as long as to subequal to the adjacent lemmas, distinctly keeled;

lower glumes 3-veined;

upper glumes 4-4.5(7+) mm, 3-veined;

calluses usually with a crown of hairs, sometimes glabrous or diffusely webbed;

lemmas 5-7.5 mm, lanceolate, 5-veined, distinctly keeled, keels, marginal veins, and sometimes the lateral veins short- to long-villous or softly puberulent, rarely glabrous, lateral veins moderately prominent, intercostal regions smooth, glabrous, margins glabrous, apices acute;

palea keels scabrous to pectinate-ciliate, intercostal regions glabrous;

anthers vestigial (0.1-0.2 mm) or (2)2.5-3.5 (4) mm.

lanceolate, distinctly shorter than the adjacent lemmas, scabrous distally, keels indistinct, scabrous, obtuse to acute;

lower glumes 3-veined;

calluses glabrous or with a crown of hairs, hairs to 0.3 mm;

lemmas 3-4.2 mm, lanceolate, weakly keeled, keels, veins, and proximal 2/3 of the intercostal regions puberulent, lateral veins obscure, margins strongly inrolled below, broadly scarious above, glabrous, apices obtuse to acute;

palea keels scabrous, sometimes hairy at midlength;

anthers 1.6-2.1 mm.

2n

= 28.

= 42.

Poa douglasii

Poa tenerrima

Distribution
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[BONAP county map]
Discussion

Poa douglasii is a dioecious endemic that grows on coastal sand dunes in California, a habitat that is being invaded by exotic species. It is rare north of Mendocino. Its hairy rachises distinguish P. douglasii from all other species of Poa in the Flora region. It differs from P. macrantha (see next), which occupies similar habitats, in this and in its usually longer glumes and lemmas.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Poa tenerrima is a rare species, endemic to serpentine barrens along the western base of the Sierra Nevada. It differs from P. secunda subsp. secunda (see previous) in combining consistently wide, open panicles, very scabrous branches, short-truncate ligules, and very fine foliage. A series of small, delicate, open-panicled plants from the California Coast Ranges, formerly included in P. tenerrima by Soreng (1993), differ in having smooth branches and longer ligules, and are better referred to P. secunda subsp. secunda. No intergradation is evident where the two taxa grow together. Plants from the Columbia River Gorge in Oregon, including the type of P. multnomae Piper, approach P. tenerrima, but are presumed to be derived from hybridization between P. nervosa (p. 545) and P. secunda (p. 586).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 24, p. 551. FNA vol. 24, p. 588.
Parent taxa Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Madropoa > subsect. Madropoa Poaceae > subfam. Pooideae > tribe Poeae > Poa > subg. Poa > sect. Secundae > subsect. Secundae
Sibling taxa
P. abbreviata, P. alpina, P. alsodes, P. ammophila, P. annua, P. arachnifera, P. arctica, P. arida, P. arnowiae, P. atropurpurea, P. autumnalis, P. bigelovii, P. bolanderi, P. bulbosa, P. chaixii, P. chambersii, P. chapmaniana, P. compressa, P. confinis, P. curtifolia, P. cusickii, P. cuspidata, P. diaboli, P. eminens, P. fendleriana, P. glauca, P. hartzii, P. howellii, P. infirma, P. interior, P. keckii, P. kelloggii, P. laxa, P. laxa × glauca, P. laxiflora, P. leibergii, P. leptocoma, P. lettermanii, P. macrantha, P. macrocalyx, P. marcida, P. napensis, P. nemoralis, P. nervosa, P. occidentalis, P. paludigena, P. palustris, P. paucispicula, P. piperi, P. porsildii, P. pratensis, P. pringlei, P. pseudoabbreviata, P. reflexa, P. rhizomata, P. saltuensis, P. secunda, P. sierrae, P. stebbinsii, P. stenantha, P. strictiramea, P. sublanata, P. suksdorfii, P. supina, P. sylvestris, P. tenerrima, P. tracyi, P. trivialis, P. unilateralis, P. wheeleri, P. wolfii, P. ×gaspensis, P. ×limosa, P. ×nematophylla
P. abbreviata, P. alpina, P. alsodes, P. ammophila, P. annua, P. arachnifera, P. arctica, P. arida, P. arnowiae, P. atropurpurea, P. autumnalis, P. bigelovii, P. bolanderi, P. bulbosa, P. chaixii, P. chambersii, P. chapmaniana, P. compressa, P. confinis, P. curtifolia, P. cusickii, P. cuspidata, P. diaboli, P. douglasii, P. eminens, P. fendleriana, P. glauca, P. hartzii, P. howellii, P. infirma, P. interior, P. keckii, P. kelloggii, P. laxa, P. laxa × glauca, P. laxiflora, P. leibergii, P. leptocoma, P. lettermanii, P. macrantha, P. macrocalyx, P. marcida, P. napensis, P. nemoralis, P. nervosa, P. occidentalis, P. paludigena, P. palustris, P. paucispicula, P. piperi, P. porsildii, P. pratensis, P. pringlei, P. pseudoabbreviata, P. reflexa, P. rhizomata, P. saltuensis, P. secunda, P. sierrae, P. stebbinsii, P. stenantha, P. strictiramea, P. sublanata, P. suksdorfii, P. supina, P. sylvestris, P. tracyi, P. trivialis, P. unilateralis, P. wheeleri, P. wolfii, P. ×gaspensis, P. ×limosa, P. ×nematophylla
Name authority Nees Scribn.
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